Chapter 11

1.True centre of sensation? Place of the mind's impressions? What is it convenient to say? What further is stated?1. Production of Sensations.—We have already seen that the true centre of sensation is some organ within the skull, probably among the gray masses at the base of the brain; but the mind never perceives impressions at that point; but, on the contrary, always refers them to the external organs of sensation. Hence, it is convenient to say, that those outer parts possess the property of sensibility. For instance, we say that we hear with the ear, taste with the tongue, and feel with the fingers. That this is not the exact truth is proven by the fact, that whenever the nerve connecting one of these organs with the brain is severed, it at once loses its capacity for sensation.2.Consciousness? During sleep? In profound insensibility?2.Consciousness, another faculty of the brain, is necessary to complete a sensation. During sleep, and in other unconscious states, the usual impressions are presented tothe ear, the nose, and the skin, but they fail to excite sensations, because the nerve-centres are inactive. In profound insensibility, from chloroform or ether, a limb may be removed without occasioning the least feeling.3.Sensibility in animals? In the earth-worm? In man?3. Variety of Sensations.—All animals have some degree of sensibility. It is of course feeble and indistinct in the lower forms of life, but increases in power and variety as we ascend the scale. In the earth-worm, the nervous system is very simple, the sensibility being moderate and alike in all parts: hence, if its body be cut into two pieces, each piece will have the same degree of feeling as before. As we approach man, however, the sensations multiply and become more acute; the organs are more complex, and special parts are endowed with special gifts. These special organs cannot be separated from the rest of the body without the loss of the functions they are designed to exercise.4.The lowest form of sensation? The highest? Sensations, how modified? What further can you state as to habitual impressions?4.The lowest form of sensation, that of simple contact, is possessed by the lowest of the animal creation. The highest forms are those by which we are enabled to know the properties of external objects, such as shape, size, sound, and color. A variety of means of communicating with the outer world is the necessary possession of a high intelligence. Sensations are modified by use. They become more acute and powerful by moderate exercise; or, they are dulled by undue excitement. The former is shown by the acute hearing of the Indian, by the sharp sight of the sailor, and by the delicate touch of the blind. The latter is exemplified by the impaired hearing of the boiler-maker, and the depraved taste of him who uses pungent condiments with his food. Again, impressions habitually presented may not be consciously felt; as is the case with the rumbling of carriages in a neighboringstreet, or the regular ticking of a clock. All sensations become less vivid with the advance of age, especially hearing and vision.5.General sensibility? What have we seen as regards the brain? Of what other structures is the same true?5. General Sensibility.—There is a property possessed by nearly all parts of the human body which we call general sensibility. We have recently seen that the brain is wholly insensitive, and may be cut or pinched without pain. The same is true of the nails, hair, the scarf-skin or external covering of the body, and a few other structures. In these parts no nerves are found. On the other hand, the sensibility of the true skin, and of mucous membranes, as of the eye and nose, is exquisite, these organs having a large supply of sensory nerve-fibres. The bones and tendons have less of these fibres, and are only moderately sensitive.6.The cause of sensibility? Painful part in a surgical operation? Benumbing the surface? How done by ether?6.The sensibility of any part of the body, then, depends upon the number of nerves present; and, as a rule, the nervous supply is proportional to the importance of the part, and to its liability to injury. When, therefore, a surgical operation is performed, the most painful part of it is the incision through the skin; the muscles, cartilage, and bone being comparatively without sensation. Hence, if we could benumb the surface, certain of the lesser operations might be undergone without great inconvenience. This is, in fact, very successfully accomplished by means of the cold produced by throwing a spray of ether, or of some other rapidly evaporating liquid, upon the part to be cut.7.Tickling? Internal sensations? The nerves of general sensibility?7.Tickling is a modification of general sensibility. At first, it excites a pleasurable sensation, but this soon passes into pain. It is only present in those parts where the sense of touch is feeble. But all impressions are not received from without: there are, also, certain internalsensations, as they are called, which depend upon the condition of the internal organs, such as appetite, hunger, thirst, the sense of satisfaction after taking food, dizziness when looking down from some lofty position, lassitude, drowsiness, fatigue, and other feelings of comfort or discomfort. General sensibility, whether of the internal or external organs of the body, chiefly depends upon the sensory fibres of the spinal nerve. The face, however, is supplied by the sensory cranial nerves. The sympathetic system has a low grade of feeling in health; but disease in the parts served by it arouses an intense degree of pain.8.Connection between pain and sensibility?8. The Sensation of Pain.—What then ispain? Is it identical with ordinary sensibility? There seems to be some necessary connection between the two feelings, for they take place through the same channels, and they are alike intense in the same situations. But sensibility habitually contributes to our sources of pleasure, the very opposite of pain; hence, these feelings cannot be identical.9.Explain the difference between pain and sensibility.9.Pain must, therefore, be a modification of the general sensibility, which follows an excessive degree of excitement of the nerves; there being a natural limit to the amount of stimulation which they will sustain. So long as this limit is observed, the part excited may be said to be simply sensitive; but when it is exceeded, the impression becomes painful. This difference between sensibility and pain is well shown by the effects of sunlight upon the eye. The indirect illumination of the sun arouses only the former feeling, and is indispensable to our comfort and existence; while the direct ray received into the eye occasions great pain.10.Dread of pain? How may its value be appreciated? Example.10. The Uses of Pain.—The dread of pain is a valuable monitor to the body. It puts us on our guard in the presence of danger; teaches moderation in the use of ourpowers; indicates the approach of disease; and calls attention to it when present. The word disease, in fact, according to its original use, had reference simply to the pain, or want of ease, which commonly attends disordered health. When we observe the serious mishaps which occur when sensibility and pain are absent, we cannot fail to appreciate its value. For example, a paralytic in taking a foot-bath, forgets to test its temperature, and putting his limbs into water while it is too hot, is severely scalded without knowing it.11.The case of the traveller? Grain of sand? The sun and child?11.A traveller, overcome by cold and fatigue, lies down and falls asleep near a large fire, and when he is aroused in the morning, it is discovered that one of his feet has been insensibly destroyed. A grain of sand, lodging in an insensitive eye, may cause inflammation and even the loss of sight. If intense light were not painful to the eye, many a child would innocently gaze upon the glories of the sun to the ruin of his sight.12.Mission of pain? Painful impressions compared with those of pleasure?12.Pain is, indeed, a present evil, but its relations with the future prove its mission merciful. Painful impressions cannot be recollected from past experience; and they cannot be called into existence by the fancy. Considered in the light of results, pain has a use above that of pleasure; for while the immoderate pursuit of the latter leads to harm, the tendency of pain is to restrict the hurtful courses of life, and in this manner to protect the body.13.What does Magendie say of the relation of pain to pleasure?13.The relations of pain to pleasure are thus described by the eminent physiologist, Magendie:—"By these sensations Nature induces us to concur in the order which she has established among organized beings. Though it may appear like sophistry to say that pain is the shadow of pleasure, yet it is certain that those who have exhausted the ordinary sources of pleasure have recourse to thecauses of pain, and gratify themselves by their effects. Do we not see in all large cities, that men who are debauched and depraved find agreeable sensations, where others experience only intolerable pain?"14.The law of Nature as regards painful sensations among animals?14.As to painful sensation among the inferior animals, the plan of Nature seems to be, that the higher the intelligence of the creature, and the more complete its power of defence, the more acute is its sensibility. We infer, therefore, that animals low in the scale of existence, and helpless, are not very liable to suffer pain.15.The sensation of contact and pain? Special sensations of man? How regarded?15. Special Sensation.—The sensations of simple contact and pain are felt by nearly all parts of the system, whether external or internal, and are the necessary consequence of the general sensibility; but, so far as the objects which surround us are concerned, these impressions are vague and passive in character, and inform the mind of none of the properties or powers of these objects. Besides these feelings, therefore, man is endowed with certain special sensations, which are positive and distinct in character, and which he can call into exercise at will, and employ in the pursuit of knowledge. For reasons relating to the original constitution of the body, these sensations are to be regarded as modifications of the general sensibility already alluded to, constructed with special reference to the different forces of Nature, of which we have any knowledge, such as heat, motion, gravity, sunlight, and the like.16.What are the special senses? Special organs for them?16.These distinct and active faculties are termed the special senses, and are five in number, viz., Touch, Taste, Smell, Sight, and Hearing. For the exercise of these senses, special organs are furnished, such as the hand, the tongue, the nose, the eye, and the ear. The manner in which the nerves of special sense terminate, varies in thecase of each organ, so that each is adapted to one set of sensations alone, and is incapable of perceiving any other. Thus the nerve of hearing is excited by the undulations of sound, and not by those of light, while the reverse is true of the nerve of sight; and the nerve of smell can appreciate neither of them, being capable only of taking cognizance of the odorous properties of bodies.17.What is said in relation to one more than the five senses?17.By some writers six senses are accorded to man; the additional one being either the sense of temperature, for as we shall presently see this is not the same as touch; or according to others, the muscular sense by which we are enabled to estimate the weights of bodies. The latter also differs in some respects from the sense of touch.18.The sense of touch, how prevalent? What is said of the hand?18. Organs of Touch.—The sense of touch is possessed by nearly all portions of the general surface of the body, but it finds its highest development in the hands. The human hand is properly regarded as the model organ of touch. The minute structure of the skin fits it admirably for this form of sensation: the cuticle, or scarf-skin, is fine and flexible, while the cutis, or true skin, contains multitudes of nerve-filaments, arranged in rows ofpapillæ, or cone-like projections, about one-hundredth of an inch in length. It is estimated that there are 20,000 of these papillæ in a square inch of the palmar surface of the hand. Now, although the nerves of the cutis are the instruments by which impressions are received and transmitted to the brain, yet the cuticle is essential to the sensation of touch. This is shown by the fact that whenever the true skin is laid bare, as by a burn or blister, the only feeling that it experiences from contact is one of pain, not that of touch.19.Office of the cuticle? Tips of the fingers? The fingers with thumb?19.The office of the cuticle is thus made evident: it is to shield the nerve filaments from direct contact withexternal objects. At the tips of the fingers, where touch is most delicate, the skin rests upon a cushion of elastic material, and receives firmness and permanence of shape by means of the nail placed upon the less sensitive side. Besides these favorable conditions, the form of the arm is such, and its motions are so easy and varied, that we are able to apply the test of touch in a great number of directions. The slender, tapering fingers, with their pliant joints, together with the strong opposable thumb, enable the hand to mold itself upon and grasp a great variety of objects; so that great as are the delicacy and grace of the hand, it is not wanting in the elements of power.20.What special importance is attributed to the hand?20.Its beauty and adaptation to the wants of man have made the hand an attractive theme for philosophers. They do not, however, always agree in their conclusions. One has the opinion that man has acquired his intelligence and achieved his place as "lord of creation," because he has this organ. Buffon, in effect, declares that with fingers twice as numerous and twice as long, we would become proportionally wiser; but Galen long ago took a more reasonable view, when he taught that "man is the wisest of animals, not because he possesses the hand; but because he is the wisest and understands its use, the hand has been given to him; for his mind, not his hand has taught him the arts." Another has well said, that "no one can study carefully the human hand and fail to be convinced of the existence of the Deity."21.The simplicity of touch? What does it teach us?21. The Sense of Touch.—Touch is the simplest of the senses. It is that which the child first calls into exercise in solving the early problems of existence; and it is that which is in the most constant use throughout life. We are brought by the touch into the most intimate relations with external objects, and by it we learn the greater number, if not the most important, of the properties ofthese objects; such as size, figure, solidity, motion, and smoothness or roughness of surface.22.Importance of the sense of touch to the development of the other senses?22.The sense of touch assists the other senses, especially that of sight, giving foundation and reality to their perceptions. Without it, the impressions received by the eye would be as vague and unreal as the figures that float through our dreams. A boy who had been blind from birth, at the age of twelve years received sight by means of a surgical operation: at first, he was unable to distinguish between a globe and a circular card, of the same color, before he had touched them. After that, he at once recognized the difference in their form. He knew the peculiarities of a dog and a cat by feeling, but not by sight, until one day, happening to take up the cat, he recognized the connection of the two sorts of impressions, those of touch and sight; and then, putting the cat down, he said: "So, puss, I shall know you next time."Fig. 45.--Sense of touch.Fig. 45.23.Liability of touch to err? Describe the illustration.23.Touch is considered the least liable to error of all the senses; yet, if that part of the skin by which the sense is exercised is removed from its customary position, a false impression may be created in the mind. This is well illustrated by an experiment, which dates from the time of Aristotle. If we cross the middle finger behind theforefinger, and then roll a marble, or some small object, upon the tips of the fingers (see Fig. 45), the impression will be that two marbles are felt. If the fingers, thus transposed, be applied to the end of the tongue, two tongues will be felt. When the nose is accidentally destroyed, the surgeon sometimes performs an operation for the purpose of forming a new one, by transplanting a partially removed piece of the skin of the forehead upon the injured part: then, if the new nose be touched or pinched, the feeling is referred to the forehead. This fact illustrates one important truth, that the nerves will re-unite after they have been cut, and feeling will be restored: if it were otherwise, a succession of slight cuts upon the fingers would seriously impair their tactile sensibility.24.The delicacy of touch? Experiments with a pair of compasses?24. The Delicacy of Touch.—Although the hand is the proper organ of this sense, yet it is exercised by various parts of the body, their degree of sensibility being proportional to the number of papillæ they contain. The varying degrees of tactile delicacy of the different parts of the surface have been measured, in an ingenious manner, by means of a pair of compasses, tipped with small pieces of cork. The two points of the compasses are touched at the same moment to the skin, the eyes being closed, and it is found that, in sensitive parts, the distance between the points may be quite slight, and yet each be plainly felt; while, in less sensitive parts, the points of the compasses are felt as a single point, although they are separated one or two inches.25.Further experiments and results?25.At the tips of the fingers, the distance between the points being one-twentieth of an inch, a double impression is felt. The distance must be twice as great, for the palm; four times as great, for the lips; and, on the forehead, it must be twenty times greater. At the middle of the back, where the touch is least acute, the points must beseparated more than two inches before they can be separately felt. Therefore, the sense of touch in the fingers is said to be fifty times more delicate than upon the posterior surface of the body.26.Exquisite delicacy of touch? The same among the blind?26.Exquisite delicacy of touch is attained by practice. This is shown in many of the lighter and more graceful employments of daily life. Without it, the skill of the painter, sculptor, and musician would be rude indeed. By training, also, the physician acquires thetactus eruditus, or discriminating touch; but among the blind, delicacy of touch is most remarkable, and it here finds its highest value; for its possession, in a measure, compensates the loss of sight by enabling them to read, by means of raised letters, to work with certain tools, and even to play upon musical instruments. A person born without sight, and without hearing or voice, may, by the education of the touch, be rescued from apparent imbecility, and be taught not only to read and write, but even to perform household and other useful labors.27.Rival candidates for the sixth sense? Give the two reasons on the subject.27. Sensations of Temperature and Weight.—Each of these sensations has been described by the physiologists as a special sense, and they are rival candidates, so to speak, for the position and title of the sixth sense. In the sensation of temperature, or the thermal sense, touch bears a part, but the two feelings appear to be distinct. In proof of this, we observe, firstly, that they are not alike intense in the same situations; as, for example, the skin of the face and elbow, where the sense of touch is feeble, is very sensitive to impressions of heat and cold. Secondly, the ability to recognize temperature may be lost by paralysis, while the sensibility of touch remains unaffected. When the skin comes in contact with a very hot substance, the sensation felt is that of pain, not of touch. In like manner, avery cold substance causes pain, not the feeling of cold. So that a red-hot iron, and solid carbonic acid (the temperature of which is 108° below zero), feel alike; and each, if pressed slightly, will produce a blister.28.The muscular sense? State what is said to illustrate the subject.28.Themuscular sense, by some considered distinct from touch, gives rise to the sensations of weight, and other forms of external resistance. That this feeling exists, is shown by the following simple experiment. If the hand be placed flat upon a table, and a somewhat heavy weight be put into it, touch alone is exercised and a feeling of pressure results; but if the hand be raised, a certain amount of muscular effort must be put forth, and thus the sensation of weight is recognized. Through the muscular sense, precision of effort is rendered possible; for by it we learn to adjust the force exerted to the weight of the object to be lifted, moved, or carried. Without it, all our movements would necessarily become ill-regulated and spasmodic. In cases of disease, where the sensibility of the lower limbs is lost, while power of motion remains, the patient is able to stand erect so long as he can see his limbs; but just as soon as his eyes are closed, he begins to waver, and will fall unless supported.29.The organ of taste? The tongue? Its powers of motion?29. The Organ of Taste.—Thetongueis the special organ of the sense of taste; but the back part of the mouth also possesses this faculty. The tongue is a muscular organ, the muscles composing it being so numerous and interwoven as to give it the freedom and variety of motion which it possesses. It can curve itself upward or downward; it can extend or contract itself; and, with its point, can sweep the cavity of the mouth, in all directions, in the search for scattered particles of food.30.Peculiarities of the tongue? Uses of the papillæ?30.The upper surface of the tongue is peculiar, being marked by the presence of innumerablepapillæ, some ofwhich are of microscopic size, resembling those that abound in the fingers, and in other parts of the body that have the sense of touch. Others are much larger, and give to the tongue its roughness of feeling and appearance. Through the medium of these papillæ, the tongue receives impressions of touch and temperature, as well as taste: indeed, its extremity is fully as delicate, in respect to tactile sensations, as the tips of the fingers themselves. It can recognize the two points of the compasses when separated not more than one-twenty-fourth of an inch; the back of it is much less sensitive to touch, while at the same time it is more highly sensitive to impressions of taste.31.Resemblance to the parts of the tongue? Powers and functions of the parts?31.Each lateral half of the tongue resembles the other in structure, and each receives the same number of nerves—three. One of these regulates motion, the other two are nerves of special sense. One of the latter supplies the front half of the tongue, and is called thegustatorynerve. This is a branch of the great cranial nerve, called the "fifth pair," which ramifies in all parts of the face. The back of the tongue is endowed with the power of taste through a nerve known as theglosso-pharyngeal, because it is distributed both to the tongue and throat. This difference in the nervous supply of the tongue becomes significant, when we learn, as we shall presently, that each part of it perceives a different class of flavors.32.Taste? What are the requisites to taste?32. The Sense of Taste.—Taste is the special sense by means of which we discover the savors, or flavoring properties of the substances, which come in contact with the tongue. Mere contact with the surface of the tongue, however, is not sufficient, but contact with the extremities of the nerves of taste within the papillæ is required. In order that the substance to be tasted may penetrate thecells covering the nerves, it must either be liquid in form, or readily soluble in the watery secretion of the mouth, the saliva. The tongue must be moist also. If the substance be insoluble, as glass or sand, or the tongue dry, the sense of taste is not awakened. In sickness, when the tongue is heavily coated, the taste is very defective, or, as is frequently expressed, "nothing tastes aright."33.Portions of the tongue endowed with taste? Where else does the sense lodge? What is stated in respect to sweet and bitter flavors? Reflex effects mentioned?33.All portions of the tongue are not alike endowed with the sense of taste, that function being limited to the posterior third, and to the margin and tip of this organ. The soft palate, also, possesses the sense of taste; hence, an article that has an agreeable flavor may very properly be spoken of as palatable, as is often done. All parts of the tongue do not perceive equally well the same flavors. Thus, the front extremity and margin, which is the portion supplied by the "fifth pair" of nerves, perceives more acutely sweet and sour tastes; but the base of the tongue, supplied by theglosso-pharyngealnerve, is especially sensitive to salt and bitter substances. The nerve of the front part of the tongue, as before stated, is in active sympathy with those of the face, while the relations of the other nerve are chiefly with the throat and stomach; so that when an intensely sour taste is perceived, the countenance is involuntarily distorted, and is said to wear an acid expression. On the other hand, a very bitter taste affects certain internal organs, and occasions a sensation of nausea, or sickness of the stomach.34.What is stated of the relations of taste with other senses?34. Relations of Taste with other Senses.—Taste is not a simple sense. Certain other sensations, as those of touch, temperature, smell, and pain, are blended and confused with it; and certain so-called tastes are really sensations of another kind. Thus an astringent taste, like that of alum, is more properly an astringent feeling, andresults from an impression made upon the nerves of touch, that ramify in the tongue. In like manner, the qualities known as smooth, oily, watery, and mealy tastes, are dependent upon these same nerves of touch. A burning or pungent taste is a sensation of pain, having its seat in the tongue and throat. A cooling taste, like that of mint, pertains to that modification of touch called the sense of temperature.35.Its dependence on smell? on sight?35.Taste is largely dependent upon the sense of smell. A considerable number of substances, like vanilla, coffee, and garlic, which appear to possess a strong and distinct flavor, have in reality a powerful odor, but only a feeble taste. When the sense of smell is interfered with by holding the nose, it becomes difficult to distinguish between substances of this class. The same effect is frequently observed when smell is blunted during an ordinary cold in the head. Sight also contributes to taste. With the eyes closed, food appears comparatively insipid; and a person smoking tobacco in the dark is unable to determine by the taste whether his cigar is lighted or not. Accordingly, it is not a bad plan to close the nose and shut the eyes when about to swallow some disagreeable medicine.36.The chief use of the sense of taste? The position of its organs? The rule as regards wholesome and unwholesome food? Remarks respecting the rule?36. Influence of Education on the Taste.—The chief use of the sense of taste appears to be to act as a guide in the selection of proper food. Hence its organs are properly placed at the entrance of the digestive canal. As a general rule, those articles which gratify the taste are wholesome; while the opposite is true of those which impress it disagreeably. This statement is more exact in reference to the early years of life than to later years, when, by reason of mischievous habits, the sense of taste has become dulled or perverted. The desires of a child are simple; he is fully satisfied with plain and wholesome articles of diet, and must usually "learn to like" thosewhich have a strongly marked flavor. Accordingly, it is far easier at this age to encourage the preference for plain food, and thus establish healthful habits, than later in life to uproot habits of indulgence in stimulating substances, after their ill effects begin to manifest themselves.37.Diversity in tastes of men? How shown? The education of the sense of taste?37.The tastes of men present the most singular diversities, partly the result of necessity and partly of habit or education. The Esquimaux like the rank smell of whale oil, which is a kind of food admirably suited to the requirements of their icy climate; and travellers who go from our climate to theirs are not slow to develop a liking for the same articles that the natives themselves enjoy. The sense of taste is rendered very acute by education, as is shown in an especial manner by those who become professional "tasters" of tea and wine.38.Location of the sense of smell? The nose? "Roof of the mouth?"38. The Sense of Smell—the Nasal Cavities.—The sense of smell is located in the delicate mucous membrane which lines the interior of the nose. That prominent feature of the face, the nose, which is merely the front boundary of the true nasal organ, is composed partly of bone and partly of cartilage. The upper part of it is united with the skull by means of a few small bones; to which circumstance is due its permanence of shape. The lower portion, or tip of the nose, contains several thin pieces of cartilage, which render it flexible and better able to resist the effects of blows and pressure. Behind the nose we find quite a spacious chamber, separated from the mouth by the hard palate, forming the "roof of the mouth," and by the soft palate (see Fig. 46); and divided into two cavities by a central partition running from before backward.39.Cavities of the nose? Obstruction of the passage of air through them?39.These nasal cavities, constituting the true beginning of the air-passages, extend from the nose backward to theupper opening of the throat, and rise as high as the junction of the nose with the forehead. The inner wall of each cavity is straight and smooth; but from the outer wall there jut into each cavity three small scroll-like bones. The structure of these bones is very light, and hence they have been called the "spongy" bones of the nose. In this manner, while the extent of surface is greatly increased by the formation of these winding passages, the cavities are rendered extremely narrow; so much so, in fact, that a moderate swelling of the mucous membrane which lines them, as from a cold, is sufficient to obstruct the passage of air through them.Fig. 46.--Section of the Right Nasal Cavity.Fig. 46.—Section of the Right Nasal Cavity.40.The special nerve of smell? Its location?40. The Nerve of Smell.—The internal surface of the nasal passages is covered by a delicate and sensitive mucous membrane. Its surface is quite extensive, following as it does, all the inequalities produced by the curved spongy bones of the nose. The upper portion of it alone is the seat of smell, since that part alone receives branches from the "first pair" of cranial nerves, or the olfactory nerve, which is the special nerve of smell (see Fig. 43). In Fig. 46 is shown the distribution of this nerve, in the form of an intricate network upon the two upper spongy bones. The nerve itself (1) does not issue from the skull, but rests upon a thin bone which separates it from the cavity of the nose; and the branches which proceed from it pass through this bone by means of numerous small openings. Theengraving represents the outer surface of the right nasal cavity; the three wave-like inequalities, upon which the nervous network is spread out, are due to the spongy bones. The left cavity is supplied in the same manner.41.Branches of the "fifth pair" of nerves? Nasal mucus? Birds?41.The nerves which ramify over the lower part of the membrane, and which endow it with sensibility to touch and pain, are branches of the "fifth pair" of nerves. An irritation applied to the parts where this nerve is distributed occasions sneezing, that is, a spasmodic contraction of the diaphragm; the object of which is the expulsion of the irritating cause. The manner in which the olfactory nerve-fibres terminate is peculiar. Unlike the extremities of other nerves, which are covered in by a greater or less thickness of tissue, these come directly to the surface of the mucous membrane, and thus come into very close contact with the odorous particles that are carried along by the respired air. The surface is at all times kept in a moist condition by an abundant flow of nasal mucus; otherwise it would become dry, hard, and insensitive from the continual passage of air to and fro in breathing. Birds, which respire more actively than men, have a special gland, for secreting a lubricating fluid, located in the air-passages of the head.42.Smell? Touch? Taste? Design of smell? Invisible and gaseous particles? The extreme fineness of the particles? Musk? In other cases?42. The Uses of the Sense of Smell.—Smell is the special sense which enables us to appreciate odors. Touch, as we have seen, is largely concerned with solid bodies; and taste, with fluids, or with solids in solution. Smell, on the other hand, is designed to afford us information in reference to substances in a volatile or gaseous form. Invisible and subtile particles emanate from odorous bodies, and are brought by the respired air in contact with the terminal filaments of the olfactory nerve, upon which an agreeable or disagreeable impression is produced.The fineness of the particles that constitute odors is often so extreme, that they elude all attempts to measure or weigh them. A piece of musk, for instance, may be kept for several years, constantly emitting perfume, without any appreciable loss of weight. In other cases, a loss of substance is perceptible, such as the essential oils, which enter into the composition of the ordinary perfumes.43.Aid given by smell? The highest use of the sense? Explain the manner.43.Smell, like taste, aids us in the choice of proper food, leading us to reject such articles as have a rank or putrid odor, and which are, as a rule, unfit to be eaten. The highest usefulness of this sense, however, consists in the protection it affords to the organs of respiration. Stationed at the gateways of the air-passages, it examines the current of air as it enters, and warns us of the presence of noxious gases, and of other and generally invisible enemies to health. Not all dangerous vapors are offensive, but almost all offensive vapors are unfit to be breathed. A number of small stiff hairs grow from the margin of the nostrils to prevent the entrance of dust and other atmospheric impurities, which would be alike injurious to the olfactory mucous membrane and to the lungs. The benevolent design of the Maker of our bodies may be observed in all parts of their mechanism; but, probably, in none is it more clearly displayed than in connection with the sense of smell.44.Sense of smell in the inferior animals? How, and in what cases, illustrated?44.The sense of smell is developed in a remarkable degree in certain of the inferior animals, and is especially acute in reference to the peculiar emanations that appear to characterize the different animals. The lion and other carnivorous beasts scent their prey from a great distance; and the fox-hound is able to track the fox through thickets and over open country for many miles; while the timid, helpless herbivora, such as the deer and sheep, find in thesense of smell a means of protection against their natural enemies, of whose approach they are in this manner warned. By training this sense in the dog, and making it subservient to his use, man is able to hunt with success certain shy and very fleet animals, which otherwise he could but seldom approach. Among men, individuals differ greatly in respect to the development of this sense; and especially in certain savage tribes it is found to be extremely delicate. Humboldt states that the natives of Peru can, by this sense, distinguish in the dark between persons of different races.45.What is sight? What information does it furnish? Composite visual sensations?45. The Sense of Sight.—Sight, or Vision, is the special sense by means of which we appreciate the color, form, size, distance, and other physical properties of the objects of external nature. Primarily, this sense furnishes us with information concerning the different shades of color and the different degrees of brightness: these are the simple sensations of sight, such as the yellowness and glitter of a gold coin. In addition to these, there are composite visual sensations, produced by the joint action of the other senses and by the use of the memory and judgment; such as, in the case of the coin, its roundness, solidity, size, its distance and direction from us. So that many of our sensations, commonly considered as due to sight, are in reality the results of intellectual processes which take place instantaneously and unconsciously.46.Comparison between sight and hearing? Relative capacity of deaf and blind?46.This faculty not only has value in the practical every-day affairs of life, but it contributes so largely to the culture of the intellect and to our higher forms of pleasure, that some writers are disposed to rate it as the first and most valuable of the senses. Others, however, maintain that the sense of hearing does not yield in importance to that of sight; and they cite in support of their positionthe fact that the blind are commonly cheerful and gay, while the deaf are inclined to be morose and melancholy. In respect to the relative capacity for receiving education in the deaf and blind, it is found that the former learn more quickly, but their attainments are not profound; while the blind acquire more slowly, but are able to study more thoroughly.47.Sight, unlike the other senses? In the case of the stars?47. Light.—The Optic Nerve.—Unlike the senses previously considered—touch, taste, and smell—sight does not bring us into immediate contact with the bodies that are examined; but, by it, we perceive the existence and qualities of objects that are at a greater or less distance from us. In the case of the stars, the distance is incalculable, while the book we read is removed but a few inches. Light is the agent which gives to this sense its wide range. The nature of this mysterious force is not known, and it is not here to be discussed; since its study belongs more properly to the province of natural philosophy.48.The undulatory theory of light? What does the theory suppose?48.It is sufficient, in this connection, to state that the theory of light now generally accepted, and which best explains the facts of optics, is that known as the undulatory theory. This theory supposes that there exists an intangible, elastic medium, which fills all space, and penetrates all transparent substances, and which is thrown into exceedingly rapid undulations or waves, by the sun and every other luminous body; the undulations being propagated with extreme rapidity, and moving not less than 186,000 miles in a second.49.The sensation of light? Optic nerve?49.These waves are thought to produce in the eye the sensation of light, in the same manner as the sonorous vibrations of the air produce in the ear the sensation of sound. That part of the eye which is sensitive to these waves is the expansion of theoptic nerve. It is sensitiveto no other impression than that of light, and it is the only nerve which is acted upon by this agent. The optic nerve, also called the "second pair" of cranial nerves, is the means of communication between the eye and the brain.50.The two nerves constituting the pair of nerves?50.The two nerves constituting the pair, arise from ganglia lying at the base of the cerebrum, one of them on each side; from which points they advance to the eyes, being united together in the middle of their course in the form of the letter X (Fig. 43—2). By this union the two eyes are enabled to act harmoniously, and in some respects to serve as a double organ. And by reason of this same intimate nervous communication, when serious disease affects one eye, the fellow-eye is extremely liable to become the seat ofsympatheticinflammation; and this, if neglected, almost certainly results in hopeless blindness.

1.True centre of sensation? Place of the mind's impressions? What is it convenient to say? What further is stated?

1. Production of Sensations.—We have already seen that the true centre of sensation is some organ within the skull, probably among the gray masses at the base of the brain; but the mind never perceives impressions at that point; but, on the contrary, always refers them to the external organs of sensation. Hence, it is convenient to say, that those outer parts possess the property of sensibility. For instance, we say that we hear with the ear, taste with the tongue, and feel with the fingers. That this is not the exact truth is proven by the fact, that whenever the nerve connecting one of these organs with the brain is severed, it at once loses its capacity for sensation.

2.Consciousness? During sleep? In profound insensibility?

2.Consciousness, another faculty of the brain, is necessary to complete a sensation. During sleep, and in other unconscious states, the usual impressions are presented tothe ear, the nose, and the skin, but they fail to excite sensations, because the nerve-centres are inactive. In profound insensibility, from chloroform or ether, a limb may be removed without occasioning the least feeling.

3.Sensibility in animals? In the earth-worm? In man?

3. Variety of Sensations.—All animals have some degree of sensibility. It is of course feeble and indistinct in the lower forms of life, but increases in power and variety as we ascend the scale. In the earth-worm, the nervous system is very simple, the sensibility being moderate and alike in all parts: hence, if its body be cut into two pieces, each piece will have the same degree of feeling as before. As we approach man, however, the sensations multiply and become more acute; the organs are more complex, and special parts are endowed with special gifts. These special organs cannot be separated from the rest of the body without the loss of the functions they are designed to exercise.

4.The lowest form of sensation? The highest? Sensations, how modified? What further can you state as to habitual impressions?

4.The lowest form of sensation, that of simple contact, is possessed by the lowest of the animal creation. The highest forms are those by which we are enabled to know the properties of external objects, such as shape, size, sound, and color. A variety of means of communicating with the outer world is the necessary possession of a high intelligence. Sensations are modified by use. They become more acute and powerful by moderate exercise; or, they are dulled by undue excitement. The former is shown by the acute hearing of the Indian, by the sharp sight of the sailor, and by the delicate touch of the blind. The latter is exemplified by the impaired hearing of the boiler-maker, and the depraved taste of him who uses pungent condiments with his food. Again, impressions habitually presented may not be consciously felt; as is the case with the rumbling of carriages in a neighboringstreet, or the regular ticking of a clock. All sensations become less vivid with the advance of age, especially hearing and vision.

5.General sensibility? What have we seen as regards the brain? Of what other structures is the same true?

5. General Sensibility.—There is a property possessed by nearly all parts of the human body which we call general sensibility. We have recently seen that the brain is wholly insensitive, and may be cut or pinched without pain. The same is true of the nails, hair, the scarf-skin or external covering of the body, and a few other structures. In these parts no nerves are found. On the other hand, the sensibility of the true skin, and of mucous membranes, as of the eye and nose, is exquisite, these organs having a large supply of sensory nerve-fibres. The bones and tendons have less of these fibres, and are only moderately sensitive.

6.The cause of sensibility? Painful part in a surgical operation? Benumbing the surface? How done by ether?

6.The sensibility of any part of the body, then, depends upon the number of nerves present; and, as a rule, the nervous supply is proportional to the importance of the part, and to its liability to injury. When, therefore, a surgical operation is performed, the most painful part of it is the incision through the skin; the muscles, cartilage, and bone being comparatively without sensation. Hence, if we could benumb the surface, certain of the lesser operations might be undergone without great inconvenience. This is, in fact, very successfully accomplished by means of the cold produced by throwing a spray of ether, or of some other rapidly evaporating liquid, upon the part to be cut.

7.Tickling? Internal sensations? The nerves of general sensibility?

7.Tickling is a modification of general sensibility. At first, it excites a pleasurable sensation, but this soon passes into pain. It is only present in those parts where the sense of touch is feeble. But all impressions are not received from without: there are, also, certain internalsensations, as they are called, which depend upon the condition of the internal organs, such as appetite, hunger, thirst, the sense of satisfaction after taking food, dizziness when looking down from some lofty position, lassitude, drowsiness, fatigue, and other feelings of comfort or discomfort. General sensibility, whether of the internal or external organs of the body, chiefly depends upon the sensory fibres of the spinal nerve. The face, however, is supplied by the sensory cranial nerves. The sympathetic system has a low grade of feeling in health; but disease in the parts served by it arouses an intense degree of pain.

8.Connection between pain and sensibility?

8. The Sensation of Pain.—What then ispain? Is it identical with ordinary sensibility? There seems to be some necessary connection between the two feelings, for they take place through the same channels, and they are alike intense in the same situations. But sensibility habitually contributes to our sources of pleasure, the very opposite of pain; hence, these feelings cannot be identical.

9.Explain the difference between pain and sensibility.

9.Pain must, therefore, be a modification of the general sensibility, which follows an excessive degree of excitement of the nerves; there being a natural limit to the amount of stimulation which they will sustain. So long as this limit is observed, the part excited may be said to be simply sensitive; but when it is exceeded, the impression becomes painful. This difference between sensibility and pain is well shown by the effects of sunlight upon the eye. The indirect illumination of the sun arouses only the former feeling, and is indispensable to our comfort and existence; while the direct ray received into the eye occasions great pain.

10.Dread of pain? How may its value be appreciated? Example.

10. The Uses of Pain.—The dread of pain is a valuable monitor to the body. It puts us on our guard in the presence of danger; teaches moderation in the use of ourpowers; indicates the approach of disease; and calls attention to it when present. The word disease, in fact, according to its original use, had reference simply to the pain, or want of ease, which commonly attends disordered health. When we observe the serious mishaps which occur when sensibility and pain are absent, we cannot fail to appreciate its value. For example, a paralytic in taking a foot-bath, forgets to test its temperature, and putting his limbs into water while it is too hot, is severely scalded without knowing it.

11.The case of the traveller? Grain of sand? The sun and child?

11.A traveller, overcome by cold and fatigue, lies down and falls asleep near a large fire, and when he is aroused in the morning, it is discovered that one of his feet has been insensibly destroyed. A grain of sand, lodging in an insensitive eye, may cause inflammation and even the loss of sight. If intense light were not painful to the eye, many a child would innocently gaze upon the glories of the sun to the ruin of his sight.

12.Mission of pain? Painful impressions compared with those of pleasure?

12.Pain is, indeed, a present evil, but its relations with the future prove its mission merciful. Painful impressions cannot be recollected from past experience; and they cannot be called into existence by the fancy. Considered in the light of results, pain has a use above that of pleasure; for while the immoderate pursuit of the latter leads to harm, the tendency of pain is to restrict the hurtful courses of life, and in this manner to protect the body.

13.What does Magendie say of the relation of pain to pleasure?

13.The relations of pain to pleasure are thus described by the eminent physiologist, Magendie:—"By these sensations Nature induces us to concur in the order which she has established among organized beings. Though it may appear like sophistry to say that pain is the shadow of pleasure, yet it is certain that those who have exhausted the ordinary sources of pleasure have recourse to thecauses of pain, and gratify themselves by their effects. Do we not see in all large cities, that men who are debauched and depraved find agreeable sensations, where others experience only intolerable pain?"

14.The law of Nature as regards painful sensations among animals?

14.As to painful sensation among the inferior animals, the plan of Nature seems to be, that the higher the intelligence of the creature, and the more complete its power of defence, the more acute is its sensibility. We infer, therefore, that animals low in the scale of existence, and helpless, are not very liable to suffer pain.

15.The sensation of contact and pain? Special sensations of man? How regarded?

15. Special Sensation.—The sensations of simple contact and pain are felt by nearly all parts of the system, whether external or internal, and are the necessary consequence of the general sensibility; but, so far as the objects which surround us are concerned, these impressions are vague and passive in character, and inform the mind of none of the properties or powers of these objects. Besides these feelings, therefore, man is endowed with certain special sensations, which are positive and distinct in character, and which he can call into exercise at will, and employ in the pursuit of knowledge. For reasons relating to the original constitution of the body, these sensations are to be regarded as modifications of the general sensibility already alluded to, constructed with special reference to the different forces of Nature, of which we have any knowledge, such as heat, motion, gravity, sunlight, and the like.

16.What are the special senses? Special organs for them?

16.These distinct and active faculties are termed the special senses, and are five in number, viz., Touch, Taste, Smell, Sight, and Hearing. For the exercise of these senses, special organs are furnished, such as the hand, the tongue, the nose, the eye, and the ear. The manner in which the nerves of special sense terminate, varies in thecase of each organ, so that each is adapted to one set of sensations alone, and is incapable of perceiving any other. Thus the nerve of hearing is excited by the undulations of sound, and not by those of light, while the reverse is true of the nerve of sight; and the nerve of smell can appreciate neither of them, being capable only of taking cognizance of the odorous properties of bodies.

17.What is said in relation to one more than the five senses?

17.By some writers six senses are accorded to man; the additional one being either the sense of temperature, for as we shall presently see this is not the same as touch; or according to others, the muscular sense by which we are enabled to estimate the weights of bodies. The latter also differs in some respects from the sense of touch.

18.The sense of touch, how prevalent? What is said of the hand?

18. Organs of Touch.—The sense of touch is possessed by nearly all portions of the general surface of the body, but it finds its highest development in the hands. The human hand is properly regarded as the model organ of touch. The minute structure of the skin fits it admirably for this form of sensation: the cuticle, or scarf-skin, is fine and flexible, while the cutis, or true skin, contains multitudes of nerve-filaments, arranged in rows ofpapillæ, or cone-like projections, about one-hundredth of an inch in length. It is estimated that there are 20,000 of these papillæ in a square inch of the palmar surface of the hand. Now, although the nerves of the cutis are the instruments by which impressions are received and transmitted to the brain, yet the cuticle is essential to the sensation of touch. This is shown by the fact that whenever the true skin is laid bare, as by a burn or blister, the only feeling that it experiences from contact is one of pain, not that of touch.

19.Office of the cuticle? Tips of the fingers? The fingers with thumb?

19.The office of the cuticle is thus made evident: it is to shield the nerve filaments from direct contact withexternal objects. At the tips of the fingers, where touch is most delicate, the skin rests upon a cushion of elastic material, and receives firmness and permanence of shape by means of the nail placed upon the less sensitive side. Besides these favorable conditions, the form of the arm is such, and its motions are so easy and varied, that we are able to apply the test of touch in a great number of directions. The slender, tapering fingers, with their pliant joints, together with the strong opposable thumb, enable the hand to mold itself upon and grasp a great variety of objects; so that great as are the delicacy and grace of the hand, it is not wanting in the elements of power.

20.What special importance is attributed to the hand?

20.Its beauty and adaptation to the wants of man have made the hand an attractive theme for philosophers. They do not, however, always agree in their conclusions. One has the opinion that man has acquired his intelligence and achieved his place as "lord of creation," because he has this organ. Buffon, in effect, declares that with fingers twice as numerous and twice as long, we would become proportionally wiser; but Galen long ago took a more reasonable view, when he taught that "man is the wisest of animals, not because he possesses the hand; but because he is the wisest and understands its use, the hand has been given to him; for his mind, not his hand has taught him the arts." Another has well said, that "no one can study carefully the human hand and fail to be convinced of the existence of the Deity."

21.The simplicity of touch? What does it teach us?

21. The Sense of Touch.—Touch is the simplest of the senses. It is that which the child first calls into exercise in solving the early problems of existence; and it is that which is in the most constant use throughout life. We are brought by the touch into the most intimate relations with external objects, and by it we learn the greater number, if not the most important, of the properties ofthese objects; such as size, figure, solidity, motion, and smoothness or roughness of surface.

22.Importance of the sense of touch to the development of the other senses?

22.The sense of touch assists the other senses, especially that of sight, giving foundation and reality to their perceptions. Without it, the impressions received by the eye would be as vague and unreal as the figures that float through our dreams. A boy who had been blind from birth, at the age of twelve years received sight by means of a surgical operation: at first, he was unable to distinguish between a globe and a circular card, of the same color, before he had touched them. After that, he at once recognized the difference in their form. He knew the peculiarities of a dog and a cat by feeling, but not by sight, until one day, happening to take up the cat, he recognized the connection of the two sorts of impressions, those of touch and sight; and then, putting the cat down, he said: "So, puss, I shall know you next time."

Fig. 45.--Sense of touch.Fig. 45.

23.Liability of touch to err? Describe the illustration.

23.Touch is considered the least liable to error of all the senses; yet, if that part of the skin by which the sense is exercised is removed from its customary position, a false impression may be created in the mind. This is well illustrated by an experiment, which dates from the time of Aristotle. If we cross the middle finger behind theforefinger, and then roll a marble, or some small object, upon the tips of the fingers (see Fig. 45), the impression will be that two marbles are felt. If the fingers, thus transposed, be applied to the end of the tongue, two tongues will be felt. When the nose is accidentally destroyed, the surgeon sometimes performs an operation for the purpose of forming a new one, by transplanting a partially removed piece of the skin of the forehead upon the injured part: then, if the new nose be touched or pinched, the feeling is referred to the forehead. This fact illustrates one important truth, that the nerves will re-unite after they have been cut, and feeling will be restored: if it were otherwise, a succession of slight cuts upon the fingers would seriously impair their tactile sensibility.

24.The delicacy of touch? Experiments with a pair of compasses?

24. The Delicacy of Touch.—Although the hand is the proper organ of this sense, yet it is exercised by various parts of the body, their degree of sensibility being proportional to the number of papillæ they contain. The varying degrees of tactile delicacy of the different parts of the surface have been measured, in an ingenious manner, by means of a pair of compasses, tipped with small pieces of cork. The two points of the compasses are touched at the same moment to the skin, the eyes being closed, and it is found that, in sensitive parts, the distance between the points may be quite slight, and yet each be plainly felt; while, in less sensitive parts, the points of the compasses are felt as a single point, although they are separated one or two inches.

25.Further experiments and results?

25.At the tips of the fingers, the distance between the points being one-twentieth of an inch, a double impression is felt. The distance must be twice as great, for the palm; four times as great, for the lips; and, on the forehead, it must be twenty times greater. At the middle of the back, where the touch is least acute, the points must beseparated more than two inches before they can be separately felt. Therefore, the sense of touch in the fingers is said to be fifty times more delicate than upon the posterior surface of the body.

26.Exquisite delicacy of touch? The same among the blind?

26.Exquisite delicacy of touch is attained by practice. This is shown in many of the lighter and more graceful employments of daily life. Without it, the skill of the painter, sculptor, and musician would be rude indeed. By training, also, the physician acquires thetactus eruditus, or discriminating touch; but among the blind, delicacy of touch is most remarkable, and it here finds its highest value; for its possession, in a measure, compensates the loss of sight by enabling them to read, by means of raised letters, to work with certain tools, and even to play upon musical instruments. A person born without sight, and without hearing or voice, may, by the education of the touch, be rescued from apparent imbecility, and be taught not only to read and write, but even to perform household and other useful labors.

27.Rival candidates for the sixth sense? Give the two reasons on the subject.

27. Sensations of Temperature and Weight.—Each of these sensations has been described by the physiologists as a special sense, and they are rival candidates, so to speak, for the position and title of the sixth sense. In the sensation of temperature, or the thermal sense, touch bears a part, but the two feelings appear to be distinct. In proof of this, we observe, firstly, that they are not alike intense in the same situations; as, for example, the skin of the face and elbow, where the sense of touch is feeble, is very sensitive to impressions of heat and cold. Secondly, the ability to recognize temperature may be lost by paralysis, while the sensibility of touch remains unaffected. When the skin comes in contact with a very hot substance, the sensation felt is that of pain, not of touch. In like manner, avery cold substance causes pain, not the feeling of cold. So that a red-hot iron, and solid carbonic acid (the temperature of which is 108° below zero), feel alike; and each, if pressed slightly, will produce a blister.

28.The muscular sense? State what is said to illustrate the subject.

28.Themuscular sense, by some considered distinct from touch, gives rise to the sensations of weight, and other forms of external resistance. That this feeling exists, is shown by the following simple experiment. If the hand be placed flat upon a table, and a somewhat heavy weight be put into it, touch alone is exercised and a feeling of pressure results; but if the hand be raised, a certain amount of muscular effort must be put forth, and thus the sensation of weight is recognized. Through the muscular sense, precision of effort is rendered possible; for by it we learn to adjust the force exerted to the weight of the object to be lifted, moved, or carried. Without it, all our movements would necessarily become ill-regulated and spasmodic. In cases of disease, where the sensibility of the lower limbs is lost, while power of motion remains, the patient is able to stand erect so long as he can see his limbs; but just as soon as his eyes are closed, he begins to waver, and will fall unless supported.

29.The organ of taste? The tongue? Its powers of motion?

29. The Organ of Taste.—Thetongueis the special organ of the sense of taste; but the back part of the mouth also possesses this faculty. The tongue is a muscular organ, the muscles composing it being so numerous and interwoven as to give it the freedom and variety of motion which it possesses. It can curve itself upward or downward; it can extend or contract itself; and, with its point, can sweep the cavity of the mouth, in all directions, in the search for scattered particles of food.

30.Peculiarities of the tongue? Uses of the papillæ?

30.The upper surface of the tongue is peculiar, being marked by the presence of innumerablepapillæ, some ofwhich are of microscopic size, resembling those that abound in the fingers, and in other parts of the body that have the sense of touch. Others are much larger, and give to the tongue its roughness of feeling and appearance. Through the medium of these papillæ, the tongue receives impressions of touch and temperature, as well as taste: indeed, its extremity is fully as delicate, in respect to tactile sensations, as the tips of the fingers themselves. It can recognize the two points of the compasses when separated not more than one-twenty-fourth of an inch; the back of it is much less sensitive to touch, while at the same time it is more highly sensitive to impressions of taste.

31.Resemblance to the parts of the tongue? Powers and functions of the parts?

31.Each lateral half of the tongue resembles the other in structure, and each receives the same number of nerves—three. One of these regulates motion, the other two are nerves of special sense. One of the latter supplies the front half of the tongue, and is called thegustatorynerve. This is a branch of the great cranial nerve, called the "fifth pair," which ramifies in all parts of the face. The back of the tongue is endowed with the power of taste through a nerve known as theglosso-pharyngeal, because it is distributed both to the tongue and throat. This difference in the nervous supply of the tongue becomes significant, when we learn, as we shall presently, that each part of it perceives a different class of flavors.

32.Taste? What are the requisites to taste?

32. The Sense of Taste.—Taste is the special sense by means of which we discover the savors, or flavoring properties of the substances, which come in contact with the tongue. Mere contact with the surface of the tongue, however, is not sufficient, but contact with the extremities of the nerves of taste within the papillæ is required. In order that the substance to be tasted may penetrate thecells covering the nerves, it must either be liquid in form, or readily soluble in the watery secretion of the mouth, the saliva. The tongue must be moist also. If the substance be insoluble, as glass or sand, or the tongue dry, the sense of taste is not awakened. In sickness, when the tongue is heavily coated, the taste is very defective, or, as is frequently expressed, "nothing tastes aright."

33.Portions of the tongue endowed with taste? Where else does the sense lodge? What is stated in respect to sweet and bitter flavors? Reflex effects mentioned?

33.All portions of the tongue are not alike endowed with the sense of taste, that function being limited to the posterior third, and to the margin and tip of this organ. The soft palate, also, possesses the sense of taste; hence, an article that has an agreeable flavor may very properly be spoken of as palatable, as is often done. All parts of the tongue do not perceive equally well the same flavors. Thus, the front extremity and margin, which is the portion supplied by the "fifth pair" of nerves, perceives more acutely sweet and sour tastes; but the base of the tongue, supplied by theglosso-pharyngealnerve, is especially sensitive to salt and bitter substances. The nerve of the front part of the tongue, as before stated, is in active sympathy with those of the face, while the relations of the other nerve are chiefly with the throat and stomach; so that when an intensely sour taste is perceived, the countenance is involuntarily distorted, and is said to wear an acid expression. On the other hand, a very bitter taste affects certain internal organs, and occasions a sensation of nausea, or sickness of the stomach.

34.What is stated of the relations of taste with other senses?

34. Relations of Taste with other Senses.—Taste is not a simple sense. Certain other sensations, as those of touch, temperature, smell, and pain, are blended and confused with it; and certain so-called tastes are really sensations of another kind. Thus an astringent taste, like that of alum, is more properly an astringent feeling, andresults from an impression made upon the nerves of touch, that ramify in the tongue. In like manner, the qualities known as smooth, oily, watery, and mealy tastes, are dependent upon these same nerves of touch. A burning or pungent taste is a sensation of pain, having its seat in the tongue and throat. A cooling taste, like that of mint, pertains to that modification of touch called the sense of temperature.

35.Its dependence on smell? on sight?

35.Taste is largely dependent upon the sense of smell. A considerable number of substances, like vanilla, coffee, and garlic, which appear to possess a strong and distinct flavor, have in reality a powerful odor, but only a feeble taste. When the sense of smell is interfered with by holding the nose, it becomes difficult to distinguish between substances of this class. The same effect is frequently observed when smell is blunted during an ordinary cold in the head. Sight also contributes to taste. With the eyes closed, food appears comparatively insipid; and a person smoking tobacco in the dark is unable to determine by the taste whether his cigar is lighted or not. Accordingly, it is not a bad plan to close the nose and shut the eyes when about to swallow some disagreeable medicine.

36.The chief use of the sense of taste? The position of its organs? The rule as regards wholesome and unwholesome food? Remarks respecting the rule?

36. Influence of Education on the Taste.—The chief use of the sense of taste appears to be to act as a guide in the selection of proper food. Hence its organs are properly placed at the entrance of the digestive canal. As a general rule, those articles which gratify the taste are wholesome; while the opposite is true of those which impress it disagreeably. This statement is more exact in reference to the early years of life than to later years, when, by reason of mischievous habits, the sense of taste has become dulled or perverted. The desires of a child are simple; he is fully satisfied with plain and wholesome articles of diet, and must usually "learn to like" thosewhich have a strongly marked flavor. Accordingly, it is far easier at this age to encourage the preference for plain food, and thus establish healthful habits, than later in life to uproot habits of indulgence in stimulating substances, after their ill effects begin to manifest themselves.

37.Diversity in tastes of men? How shown? The education of the sense of taste?

37.The tastes of men present the most singular diversities, partly the result of necessity and partly of habit or education. The Esquimaux like the rank smell of whale oil, which is a kind of food admirably suited to the requirements of their icy climate; and travellers who go from our climate to theirs are not slow to develop a liking for the same articles that the natives themselves enjoy. The sense of taste is rendered very acute by education, as is shown in an especial manner by those who become professional "tasters" of tea and wine.

38.Location of the sense of smell? The nose? "Roof of the mouth?"

38. The Sense of Smell—the Nasal Cavities.—The sense of smell is located in the delicate mucous membrane which lines the interior of the nose. That prominent feature of the face, the nose, which is merely the front boundary of the true nasal organ, is composed partly of bone and partly of cartilage. The upper part of it is united with the skull by means of a few small bones; to which circumstance is due its permanence of shape. The lower portion, or tip of the nose, contains several thin pieces of cartilage, which render it flexible and better able to resist the effects of blows and pressure. Behind the nose we find quite a spacious chamber, separated from the mouth by the hard palate, forming the "roof of the mouth," and by the soft palate (see Fig. 46); and divided into two cavities by a central partition running from before backward.

39.Cavities of the nose? Obstruction of the passage of air through them?

39.These nasal cavities, constituting the true beginning of the air-passages, extend from the nose backward to theupper opening of the throat, and rise as high as the junction of the nose with the forehead. The inner wall of each cavity is straight and smooth; but from the outer wall there jut into each cavity three small scroll-like bones. The structure of these bones is very light, and hence they have been called the "spongy" bones of the nose. In this manner, while the extent of surface is greatly increased by the formation of these winding passages, the cavities are rendered extremely narrow; so much so, in fact, that a moderate swelling of the mucous membrane which lines them, as from a cold, is sufficient to obstruct the passage of air through them.

Fig. 46.--Section of the Right Nasal Cavity.Fig. 46.—Section of the Right Nasal Cavity.

40.The special nerve of smell? Its location?

40. The Nerve of Smell.—The internal surface of the nasal passages is covered by a delicate and sensitive mucous membrane. Its surface is quite extensive, following as it does, all the inequalities produced by the curved spongy bones of the nose. The upper portion of it alone is the seat of smell, since that part alone receives branches from the "first pair" of cranial nerves, or the olfactory nerve, which is the special nerve of smell (see Fig. 43). In Fig. 46 is shown the distribution of this nerve, in the form of an intricate network upon the two upper spongy bones. The nerve itself (1) does not issue from the skull, but rests upon a thin bone which separates it from the cavity of the nose; and the branches which proceed from it pass through this bone by means of numerous small openings. Theengraving represents the outer surface of the right nasal cavity; the three wave-like inequalities, upon which the nervous network is spread out, are due to the spongy bones. The left cavity is supplied in the same manner.

41.Branches of the "fifth pair" of nerves? Nasal mucus? Birds?

41.The nerves which ramify over the lower part of the membrane, and which endow it with sensibility to touch and pain, are branches of the "fifth pair" of nerves. An irritation applied to the parts where this nerve is distributed occasions sneezing, that is, a spasmodic contraction of the diaphragm; the object of which is the expulsion of the irritating cause. The manner in which the olfactory nerve-fibres terminate is peculiar. Unlike the extremities of other nerves, which are covered in by a greater or less thickness of tissue, these come directly to the surface of the mucous membrane, and thus come into very close contact with the odorous particles that are carried along by the respired air. The surface is at all times kept in a moist condition by an abundant flow of nasal mucus; otherwise it would become dry, hard, and insensitive from the continual passage of air to and fro in breathing. Birds, which respire more actively than men, have a special gland, for secreting a lubricating fluid, located in the air-passages of the head.

42.Smell? Touch? Taste? Design of smell? Invisible and gaseous particles? The extreme fineness of the particles? Musk? In other cases?

42. The Uses of the Sense of Smell.—Smell is the special sense which enables us to appreciate odors. Touch, as we have seen, is largely concerned with solid bodies; and taste, with fluids, or with solids in solution. Smell, on the other hand, is designed to afford us information in reference to substances in a volatile or gaseous form. Invisible and subtile particles emanate from odorous bodies, and are brought by the respired air in contact with the terminal filaments of the olfactory nerve, upon which an agreeable or disagreeable impression is produced.The fineness of the particles that constitute odors is often so extreme, that they elude all attempts to measure or weigh them. A piece of musk, for instance, may be kept for several years, constantly emitting perfume, without any appreciable loss of weight. In other cases, a loss of substance is perceptible, such as the essential oils, which enter into the composition of the ordinary perfumes.

43.Aid given by smell? The highest use of the sense? Explain the manner.

43.Smell, like taste, aids us in the choice of proper food, leading us to reject such articles as have a rank or putrid odor, and which are, as a rule, unfit to be eaten. The highest usefulness of this sense, however, consists in the protection it affords to the organs of respiration. Stationed at the gateways of the air-passages, it examines the current of air as it enters, and warns us of the presence of noxious gases, and of other and generally invisible enemies to health. Not all dangerous vapors are offensive, but almost all offensive vapors are unfit to be breathed. A number of small stiff hairs grow from the margin of the nostrils to prevent the entrance of dust and other atmospheric impurities, which would be alike injurious to the olfactory mucous membrane and to the lungs. The benevolent design of the Maker of our bodies may be observed in all parts of their mechanism; but, probably, in none is it more clearly displayed than in connection with the sense of smell.

44.Sense of smell in the inferior animals? How, and in what cases, illustrated?

44.The sense of smell is developed in a remarkable degree in certain of the inferior animals, and is especially acute in reference to the peculiar emanations that appear to characterize the different animals. The lion and other carnivorous beasts scent their prey from a great distance; and the fox-hound is able to track the fox through thickets and over open country for many miles; while the timid, helpless herbivora, such as the deer and sheep, find in thesense of smell a means of protection against their natural enemies, of whose approach they are in this manner warned. By training this sense in the dog, and making it subservient to his use, man is able to hunt with success certain shy and very fleet animals, which otherwise he could but seldom approach. Among men, individuals differ greatly in respect to the development of this sense; and especially in certain savage tribes it is found to be extremely delicate. Humboldt states that the natives of Peru can, by this sense, distinguish in the dark between persons of different races.

45.What is sight? What information does it furnish? Composite visual sensations?

45. The Sense of Sight.—Sight, or Vision, is the special sense by means of which we appreciate the color, form, size, distance, and other physical properties of the objects of external nature. Primarily, this sense furnishes us with information concerning the different shades of color and the different degrees of brightness: these are the simple sensations of sight, such as the yellowness and glitter of a gold coin. In addition to these, there are composite visual sensations, produced by the joint action of the other senses and by the use of the memory and judgment; such as, in the case of the coin, its roundness, solidity, size, its distance and direction from us. So that many of our sensations, commonly considered as due to sight, are in reality the results of intellectual processes which take place instantaneously and unconsciously.

46.Comparison between sight and hearing? Relative capacity of deaf and blind?

46.This faculty not only has value in the practical every-day affairs of life, but it contributes so largely to the culture of the intellect and to our higher forms of pleasure, that some writers are disposed to rate it as the first and most valuable of the senses. Others, however, maintain that the sense of hearing does not yield in importance to that of sight; and they cite in support of their positionthe fact that the blind are commonly cheerful and gay, while the deaf are inclined to be morose and melancholy. In respect to the relative capacity for receiving education in the deaf and blind, it is found that the former learn more quickly, but their attainments are not profound; while the blind acquire more slowly, but are able to study more thoroughly.

47.Sight, unlike the other senses? In the case of the stars?

47. Light.—The Optic Nerve.—Unlike the senses previously considered—touch, taste, and smell—sight does not bring us into immediate contact with the bodies that are examined; but, by it, we perceive the existence and qualities of objects that are at a greater or less distance from us. In the case of the stars, the distance is incalculable, while the book we read is removed but a few inches. Light is the agent which gives to this sense its wide range. The nature of this mysterious force is not known, and it is not here to be discussed; since its study belongs more properly to the province of natural philosophy.

48.The undulatory theory of light? What does the theory suppose?

48.It is sufficient, in this connection, to state that the theory of light now generally accepted, and which best explains the facts of optics, is that known as the undulatory theory. This theory supposes that there exists an intangible, elastic medium, which fills all space, and penetrates all transparent substances, and which is thrown into exceedingly rapid undulations or waves, by the sun and every other luminous body; the undulations being propagated with extreme rapidity, and moving not less than 186,000 miles in a second.

49.The sensation of light? Optic nerve?

49.These waves are thought to produce in the eye the sensation of light, in the same manner as the sonorous vibrations of the air produce in the ear the sensation of sound. That part of the eye which is sensitive to these waves is the expansion of theoptic nerve. It is sensitiveto no other impression than that of light, and it is the only nerve which is acted upon by this agent. The optic nerve, also called the "second pair" of cranial nerves, is the means of communication between the eye and the brain.

50.The two nerves constituting the pair of nerves?

50.The two nerves constituting the pair, arise from ganglia lying at the base of the cerebrum, one of them on each side; from which points they advance to the eyes, being united together in the middle of their course in the form of the letter X (Fig. 43—2). By this union the two eyes are enabled to act harmoniously, and in some respects to serve as a double organ. And by reason of this same intimate nervous communication, when serious disease affects one eye, the fellow-eye is extremely liable to become the seat ofsympatheticinflammation; and this, if neglected, almost certainly results in hopeless blindness.


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