Type.—Male, adult, skeleton and skin; no. 32820, Mus. Vert. Zoöl.; Half Moon Bay, San Mateo County, California; received at Museum of Vertebrate Zoölogy, May 4, 1922, through A. L. Hagedoorn, after having been in captivity a few days where death occurred owing to injuries received in trap; original no. 1590.The skull has each of the zygomatic arches and the anterior end of the nasals broken through. The only part missing is the central two millimeters of the left zygomatic arch. The teeth all are present and entire. The skeleton appears to be complete except for the bones of the feet, which are preserved within the skin. The skin is well made and in good condition.Range.—Altitudinally, sea level to more than 4000 feet; Sonoran and Transition life-zones of Coast Range and coast of California from San Francisco Bay south to Point Conception, Santa Barbara County, California. See figures29and30on pages 221 and 314.Characters for ready recognition.—Differs fromM. f. munda,xanthogenys, andpulchraby having inside of ears darker than back rather than same color as back, and fromxanthogenysandpulchrain near (l) Antique Brown color of upper parts rather than Buckthorn Brown or near (16j) Buckthorn Brown to near (h) Yellow Ocher respectively; fromM. f. latirostraby postorbital breadth, of adult males and females, less, rather than more, than width of basioccipital measured from medial margin of one foramen lacerum posterior to its opposite.Description.—Size.—Male: Five adults from Palo Alto, Santa Clara County, yield average and extreme measurements as follows: Total length, 447 (412-465); length of tail, 167 (147-175); length of hind foot, 46 (45-47). Corresponding measurements of four adults from San Francisco are: 412 (394-435); 153 (145-160); 43.5 (41-46). Corresponding measurements of five adults and subadults from Berkeley, Alameda County, are: 419 (390-448); 148 (135-160); 44 (42-47). Tail averages 59 per cent as long as head and body in series from Palo Alto and in one from San Francisco. The average of 55 for the Berkeley series probably reflects a lesser average age. Length of hind foot less than basal length. The type specimen measures, 415, 150, 43. It is smaller than the mean.Female: A subadult from Palo Alto measures: Total length, 368; length of tail, 126; length of hind foot, 39. An adult and two subadults from Berkeley measure, respectively, as follows: Total length, 347, 365, 340; length of tail, 134, 123, 125; length of hind foot, 37, 38.4, 36.5. In these four females the tail averages 55 per cent as long as head and body. Length of hind foot less than basal length.The average differences in external measurements of the two sexes, as represented by specimens from Berkeley, Alameda County, are: Total length, 68; length of tail, 21; length of hind foot, 7. Eight adult males weigh 249 (217-335) grams and one adult female 123 grams.Externals.—Longest facial vibrissae brownish like dark color of head and extending beyond ear; carpal vibrissae mostly color of underparts and extending to apical pad of fifth digit; hairiness of foot-soles slightly more than shown in figure20.Color.—Spot between eyes, band, confluent with color of underparts, on each side of head extending anterodorsally anterior to ear, and posterior third of each upper lip tinged with color of underparts or, less often, pure white; chin and lower lips white; remainder of sides and top of head posteriorly to, or a little behind, a line connecting posterior margins of ears, blackish; inside of pinna of ear, and sometimes outside of pinna, blackish; dark spot posterior to each angle of mouth present on each side in three-fourths of specimens; tip of tail black; remainder of upper parts near (l) Antique Brown, and with more yellow than tone 3 of Raw Umber of Oberthür and Dauthenay, pl. 301. Often with more blackish and red in winter. Underparts near (atoc) Ochraceous-Buff or Ochraceous-Salmon. Ochraceous-Salmon in some juveniles. Color of underparts extends distally on posterior sides of forelegs over toes onto antipalmar faces of feet and wrists, and on medial sides of hind limbs over antiplantar faces of toes. Least width of color of underparts averaging, in 17 adult males (Berkeley, 5; San Francisco, 5; Palo Alto, 7), 55 (40-73) per cent of greatest width of color of upper parts. Black tip of tail in same series of males averaging 51 (35-60) mm., thus averaging longer than hind foot and 33 per cent of length of tail (Palo Alto and San Francisco, 31 per cent; Berkeley, 35 per cent). In 8 adult females, least color of underparts amounts to 55 (47-62) per cent of greatest width of color of upper parts. Black tip of tail averages 41.5 (28-50) mm., thus averaging longer than hind foot and 32 per cent of length of tail-vertebrae.Skull and teeth.—Male (based on six adults from Stanford Univ. and vicinity): See measurements and plates22-24; weight (four adults), 5.4 (5.0-5.9) grams; basilar length, 47 (46.1-48.1); zygomatic breadth more than distance between condylar foramen and M1, or than between anterior palatine foramen and anterior margin of tympanic bulla; mastoid breadth more than postpalatal length; postorbital breadth less than length of upper premolars (less than distance between posterior borders of P4 and P2) and less than width of basioccipital measured from medial margin of one foramen lacerum posterior to its opposite; interorbital breadth not greater than distance between foramen opticum and anterior margin of tympanic bulla; breadth of rostrum less than length of tympanic bulla; least width of palate less than lateral length of P4; anterior margin of tympanic bulla as far posterior to foramen ovale as width of 3 or 4 (including I3) upper incisors; height of tympanic bulla more than distance from its anterior margin to foramen ovale; length of tympanic bulla more or less than (about equal to) length of lower molar and premolar tooth-row and longer or shorter (usually shorter) than rostrum; anterior margin of masseteric fossa below anterior half of m2.Female (based on three adults, Hayward, Palo Alto, and Morro): See measurements and plates34-36; weight (no. 43574, from Morro) 2.7 grams; basilar length, 41.2 (40.2-42.2); zygomatic breadth more or less than distance between condylar foramen and M1 and more or less than distance between anterior palatine foramen and anterior margin of tympanic bulla; postorbital breadth less than length of upper premolars and less than width of basioccipital measured from medial margin of one foramen lacerum posterior to its opposite; least width of palate less than lateral length of P4; tympanic bulla as far posterior to foramen ovale as width of 3 (including I3) upper incisors; height of tympanic bulla more than distance from its anterior margin to foramen ovale; length of tympanic bulla more than length of lower molar and premolar tooth-row and longer or shorter than rostrum.The skull of the female averages 50 per cent lighter than that of the average male.Comparisons of the skull of the male with those ofM. f. latirostra,pulchra,xanthogenys, andmundaare made in the accounts of those subspecies.
Type.—Male, adult, skeleton and skin; no. 32820, Mus. Vert. Zoöl.; Half Moon Bay, San Mateo County, California; received at Museum of Vertebrate Zoölogy, May 4, 1922, through A. L. Hagedoorn, after having been in captivity a few days where death occurred owing to injuries received in trap; original no. 1590.
The skull has each of the zygomatic arches and the anterior end of the nasals broken through. The only part missing is the central two millimeters of the left zygomatic arch. The teeth all are present and entire. The skeleton appears to be complete except for the bones of the feet, which are preserved within the skin. The skin is well made and in good condition.
Range.—Altitudinally, sea level to more than 4000 feet; Sonoran and Transition life-zones of Coast Range and coast of California from San Francisco Bay south to Point Conception, Santa Barbara County, California. See figures29and30on pages 221 and 314.
Characters for ready recognition.—Differs fromM. f. munda,xanthogenys, andpulchraby having inside of ears darker than back rather than same color as back, and fromxanthogenysandpulchrain near (l) Antique Brown color of upper parts rather than Buckthorn Brown or near (16j) Buckthorn Brown to near (h) Yellow Ocher respectively; fromM. f. latirostraby postorbital breadth, of adult males and females, less, rather than more, than width of basioccipital measured from medial margin of one foramen lacerum posterior to its opposite.
Description.—Size.—Male: Five adults from Palo Alto, Santa Clara County, yield average and extreme measurements as follows: Total length, 447 (412-465); length of tail, 167 (147-175); length of hind foot, 46 (45-47). Corresponding measurements of four adults from San Francisco are: 412 (394-435); 153 (145-160); 43.5 (41-46). Corresponding measurements of five adults and subadults from Berkeley, Alameda County, are: 419 (390-448); 148 (135-160); 44 (42-47). Tail averages 59 per cent as long as head and body in series from Palo Alto and in one from San Francisco. The average of 55 for the Berkeley series probably reflects a lesser average age. Length of hind foot less than basal length. The type specimen measures, 415, 150, 43. It is smaller than the mean.
Female: A subadult from Palo Alto measures: Total length, 368; length of tail, 126; length of hind foot, 39. An adult and two subadults from Berkeley measure, respectively, as follows: Total length, 347, 365, 340; length of tail, 134, 123, 125; length of hind foot, 37, 38.4, 36.5. In these four females the tail averages 55 per cent as long as head and body. Length of hind foot less than basal length.
The average differences in external measurements of the two sexes, as represented by specimens from Berkeley, Alameda County, are: Total length, 68; length of tail, 21; length of hind foot, 7. Eight adult males weigh 249 (217-335) grams and one adult female 123 grams.
Externals.—Longest facial vibrissae brownish like dark color of head and extending beyond ear; carpal vibrissae mostly color of underparts and extending to apical pad of fifth digit; hairiness of foot-soles slightly more than shown in figure20.
Color.—Spot between eyes, band, confluent with color of underparts, on each side of head extending anterodorsally anterior to ear, and posterior third of each upper lip tinged with color of underparts or, less often, pure white; chin and lower lips white; remainder of sides and top of head posteriorly to, or a little behind, a line connecting posterior margins of ears, blackish; inside of pinna of ear, and sometimes outside of pinna, blackish; dark spot posterior to each angle of mouth present on each side in three-fourths of specimens; tip of tail black; remainder of upper parts near (l) Antique Brown, and with more yellow than tone 3 of Raw Umber of Oberthür and Dauthenay, pl. 301. Often with more blackish and red in winter. Underparts near (atoc) Ochraceous-Buff or Ochraceous-Salmon. Ochraceous-Salmon in some juveniles. Color of underparts extends distally on posterior sides of forelegs over toes onto antipalmar faces of feet and wrists, and on medial sides of hind limbs over antiplantar faces of toes. Least width of color of underparts averaging, in 17 adult males (Berkeley, 5; San Francisco, 5; Palo Alto, 7), 55 (40-73) per cent of greatest width of color of upper parts. Black tip of tail in same series of males averaging 51 (35-60) mm., thus averaging longer than hind foot and 33 per cent of length of tail (Palo Alto and San Francisco, 31 per cent; Berkeley, 35 per cent). In 8 adult females, least color of underparts amounts to 55 (47-62) per cent of greatest width of color of upper parts. Black tip of tail averages 41.5 (28-50) mm., thus averaging longer than hind foot and 32 per cent of length of tail-vertebrae.
Skull and teeth.—Male (based on six adults from Stanford Univ. and vicinity): See measurements and plates22-24; weight (four adults), 5.4 (5.0-5.9) grams; basilar length, 47 (46.1-48.1); zygomatic breadth more than distance between condylar foramen and M1, or than between anterior palatine foramen and anterior margin of tympanic bulla; mastoid breadth more than postpalatal length; postorbital breadth less than length of upper premolars (less than distance between posterior borders of P4 and P2) and less than width of basioccipital measured from medial margin of one foramen lacerum posterior to its opposite; interorbital breadth not greater than distance between foramen opticum and anterior margin of tympanic bulla; breadth of rostrum less than length of tympanic bulla; least width of palate less than lateral length of P4; anterior margin of tympanic bulla as far posterior to foramen ovale as width of 3 or 4 (including I3) upper incisors; height of tympanic bulla more than distance from its anterior margin to foramen ovale; length of tympanic bulla more or less than (about equal to) length of lower molar and premolar tooth-row and longer or shorter (usually shorter) than rostrum; anterior margin of masseteric fossa below anterior half of m2.
Female (based on three adults, Hayward, Palo Alto, and Morro): See measurements and plates34-36; weight (no. 43574, from Morro) 2.7 grams; basilar length, 41.2 (40.2-42.2); zygomatic breadth more or less than distance between condylar foramen and M1 and more or less than distance between anterior palatine foramen and anterior margin of tympanic bulla; postorbital breadth less than length of upper premolars and less than width of basioccipital measured from medial margin of one foramen lacerum posterior to its opposite; least width of palate less than lateral length of P4; tympanic bulla as far posterior to foramen ovale as width of 3 (including I3) upper incisors; height of tympanic bulla more than distance from its anterior margin to foramen ovale; length of tympanic bulla more than length of lower molar and premolar tooth-row and longer or shorter than rostrum.
The skull of the female averages 50 per cent lighter than that of the average male.
Comparisons of the skull of the male with those ofM. f. latirostra,pulchra,xanthogenys, andmundaare made in the accounts of those subspecies.
Remarks.—LikeM. f. latirostra,nigriaurislong bore the namexanthogenys. The fairly adequate lot of specimens is divided between the collections of several institutions. The most satisfactory material in any one collection is in the Stanford University Natural History Museum where local specimens have been accumulated over a period of many years.
No actual intergrade betweennigriaurisandxanthogenyshas been seen, although the specimens from Los Banos, referred toxanthogenys, have large skulls as innigriauris. Intergradation withlatirostrais shown by specimens, referred tolatirostra, from the Los Angeles area. Also the one adult male from 5 miles southeast of Santa Margarita, San Luis Obispo County, is of small size and in this respect approacheslatirostra. The range ofnigriaurisis separated from that ofmundaby San Francisco Bay, Carquinez Straits, and I suppose by the lower part of the San Joaquin River. On the basis of color of the inside of the pinna of the ear, the two subspecies are uniformly distinct. Intergradation is assumed to occur through the subspeciesxanthogenys.
None of the 26 adult and subadult specimens examined for evidences of infestation of the frontal sinuses by parasites shows malformation of the sinuses.
Specimens examined.—Total number, 103, arranged by counties from north to south. Unless otherwise indicated specimens are in the Museum of Vertebrate Zoölogy.California.Contra Costa County: Glen Frazer Station, 1; 2 mi. W Pinole, 1[13]; 1 mi. E Pinole, 1; Richmond, 1[13]; Lafayette, 1; 7 mi. E Clayton, 1; Moraga Valley, 1; Pinehurst, Redwood Canyon, 1; Concord, 1.Alameda County: Berkeley, 11; Oakland, 1; Piedmont, 1; Haywards, 2; near Haywards, 2; 10 mi. E Haywards, 1[91]; Redwood Canyon, 1; Calaveras Dam, 1.San Francisco County: San Francisco, 11 (5[8], 2[91], 1[60], 1[7]); Ocean View, 1[68]; Visitation Valley, 1.San Mateo County: Moss Beach, 1; Half Moon Bay, 1; Redwood City, 1[87]; Menlo Park, 9 (5[87], 2[68]); no locality more definite than county, 1[8].Santa Clara County: 1/4 mi. N Milpitas, 1; 1/4 mi. S Milpitas, 1; Stanford University, 6 (4[68], 2[91]); Palo Alto, 11 (6[41], 2[60], 1[75], 1[87]).Santa Cruz County: 3 mi. E Santa Cruz, 1; 2-1/2 mi. E Santa Cruz, 1; Santa Cruz, 6 (2[91], 1[68], 1[4]).Monterey County: 1 mi. E mouth Salinas River, 10 ft., 1[37]; Pacific Grove, 1[8]; Monterey, 2 (1[7]); Carmel, 1[8]; Carmel Valley, 1[68]; Point Lobos, 1; Gonzales, 1.San Luis Obispo County: 5 mi. SE Santa Margarita, 1; Morro, 1[91]; 3-1/2 mi. S Oceano, 6.Santa Barbara County: Santa Maria, 1[87]; 5 mi. N Las Cruces, 1; 7 mi. W Gaviota, 1; Gaviota, 1.
Specimens examined.—Total number, 103, arranged by counties from north to south. Unless otherwise indicated specimens are in the Museum of Vertebrate Zoölogy.
California.Contra Costa County: Glen Frazer Station, 1; 2 mi. W Pinole, 1[13]; 1 mi. E Pinole, 1; Richmond, 1[13]; Lafayette, 1; 7 mi. E Clayton, 1; Moraga Valley, 1; Pinehurst, Redwood Canyon, 1; Concord, 1.Alameda County: Berkeley, 11; Oakland, 1; Piedmont, 1; Haywards, 2; near Haywards, 2; 10 mi. E Haywards, 1[91]; Redwood Canyon, 1; Calaveras Dam, 1.San Francisco County: San Francisco, 11 (5[8], 2[91], 1[60], 1[7]); Ocean View, 1[68]; Visitation Valley, 1.San Mateo County: Moss Beach, 1; Half Moon Bay, 1; Redwood City, 1[87]; Menlo Park, 9 (5[87], 2[68]); no locality more definite than county, 1[8].Santa Clara County: 1/4 mi. N Milpitas, 1; 1/4 mi. S Milpitas, 1; Stanford University, 6 (4[68], 2[91]); Palo Alto, 11 (6[41], 2[60], 1[75], 1[87]).Santa Cruz County: 3 mi. E Santa Cruz, 1; 2-1/2 mi. E Santa Cruz, 1; Santa Cruz, 6 (2[91], 1[68], 1[4]).Monterey County: 1 mi. E mouth Salinas River, 10 ft., 1[37]; Pacific Grove, 1[8]; Monterey, 2 (1[7]); Carmel, 1[8]; Carmel Valley, 1[68]; Point Lobos, 1; Gonzales, 1.San Luis Obispo County: 5 mi. SE Santa Margarita, 1; Morro, 1[91]; 3-1/2 mi. S Oceano, 6.Santa Barbara County: Santa Maria, 1[87]; 5 mi. N Las Cruces, 1; 7 mi. W Gaviota, 1; Gaviota, 1.
Long-tailed Weasel
Plates1,22,23,24,34,35and 36
Mustela frenata latirostraHall, Carnegie Instit. Washington Publ. 473:96, November 20, 1936.Putorius xanthogenys, Baird, Mamm. N. Amer., p. 176, 1858 (part); Stephens, California mammals, p. 246, 1906; Merriam, N. Amer. Fauna, 11:25, June 30, 1896 (part).Putorius (Gale) brasilianus frenatus, Coues, Fur-bearing animals, p. 142 (part).Mustela xanthogenys xanthogenys, Miller, U. S. Nat. Mus. Bull., 79:99, December 31, 1912; Grinnell, Univ. California Publ. Zoöl., 40:102, September 26, 1933.Mustela arizonensis, Grinnell and Swarth, Univ. California Publ. Zoöl. 10, 376, October 31, 1913.
Mustela frenata latirostraHall, Carnegie Instit. Washington Publ. 473:96, November 20, 1936.
Putorius xanthogenys, Baird, Mamm. N. Amer., p. 176, 1858 (part); Stephens, California mammals, p. 246, 1906; Merriam, N. Amer. Fauna, 11:25, June 30, 1896 (part).
Putorius (Gale) brasilianus frenatus, Coues, Fur-bearing animals, p. 142 (part).
Mustela xanthogenys xanthogenys, Miller, U. S. Nat. Mus. Bull., 79:99, December 31, 1912; Grinnell, Univ. California Publ. Zoöl., 40:102, September 26, 1933.
Mustela arizonensis, Grinnell and Swarth, Univ. California Publ. Zoöl. 10, 376, October 31, 1913.
Type.—Male, adult, skull and skin; no. 3257, Mus. Vert. Zoöl.; San Diego, San Diego County, California; May 20, 1907; obtained by Frank X. Holzner.Right M1 is missing and the part of the jaw bearing this tooth is broken away. With this exception the skull is complete and unbroken and the teeth are all present and entire. The skin is fairly well made and in good condition except that it is slightly soiled.Range.—Altitudinally sea level to 8000 feet (Tahquitz Valley, San Jacinto Mountains); Sonoran and Transition life-zones of coast and mountains west of Mohave and Imperial deserts of southern California from Point Conception and Cuyama Valley southward at least to Mexican boundary. See figures29and30on pages 221 and 314.Characters for ready recognition.—Differs fromM. f. nigriaurisby having postorbital breadth of adult males and females, more, rather than less, than width of basioccipital measured from medial margin of one foramen lacerum posterior to its opposite; fromM. f. pulchraby having tympanic bulla longer than rostrum (orbitonasal length) and by near (l) Antique Brown rather than near (16j) Buckthorn Brown to near (h) Yellow Ocher color of upper parts.Description.—Size.—Male: Six adults and subadults from San Diego yield average and extreme measurements as follows: Total length, 439 (428-449); length of tail, 153 (142-160); length of hind foot, 45 (40-47). Corresponding measurements for a series of eight adult males from the vicinity of Los Angeles are: 416 (394-428); 158 (151-166); 44 (40-47). In the series from San Diego the tail averages 54 per cent as long as head and body. In the series from Los Angeles the average is 61 per cent. Length of hind foot in each series, less than basal length. The type specimen measures, 435, 142, 42.Female: No. 5070, adult, from San Diego, measures 367, 141, 38. Nos. 22 and 6748 from Santa Ysabel, measure: 359, 380; 130, 140; 39, 35. No. 7194 from Jamacha measures, 358, 125, 35. Three adult females from Los Angeles yield the following: Total length, 373, 345, 368; length of tail, 150, 122, 134; length of hind foot,—, 41, 41. In no. 5070 the tail is 62 per cent as long as the head and body and in the three from Los Angeles it averages 60 (55-67) per cent. Length of hind foot, in each case, less than basal length.The average differences in external measurements of the two sexes as shown by the six males from San Diego and the four females from San Diego County are: Total length, 73; length of tail, 19; length of hind foot, 8. Corresponding differences shown by the eight males and three females from Los Angeles are: 54, 23, 3.Externals.—Longest facial vibrissae brownish, like dark color of head and extending beyond ear; carpal vibrissae mostly color of underparts and extending to apical pad of fifth digit; hairiness of foot-soles slightly more than shown in figure20.Color.—Spot between eyes, band confluent with color of underparts on each side of head extending anterodorsally anterior to ear, and posterior third of each upper lip tinged with color of underparts or, less often, white; chin and lower lips white; remainder of sides and top of head posteriorly to near line connecting posterior margins of ears, blackish; inside of pinna of ear, and sometimes outside of pinna, blackish; dark spot posterior to each angle of mouth present on each side in three-fourths of specimens; tip of tail black; remainder of upper parts near (l) Antique Brown, and with more yellow than tone 3 of Raw Umber of Oberthür and Dauthenay, pl. 301. Underparts Ochraceous-Buff to Warm Buff and in some specimens from Los Angeles and Ventura counties Ochraceous-Orange, especially in young and juveniles. Color of underparts extends distally on posterior sides of forelegs over toes onto antipalmar faces of feet and wrists and on medial sides of hind limbs over antiplantar faces of toes. Least width of color of underparts averaging, in 15 adult and subadult males from San Diego County, 54 (35-75) per cent of greatest width of color of upper parts. Black tip of tail in same series of males averaging 54.5 (46-60) mm. long. Thus averaging longer than hind foot and 35 per cent of length of tail-vertebrae.Skull and teeth.—Male (based on 6 adults from San Diego County). See measurements and plates22-24. As described inM. f. nigriaurisexcept that: Weight (4 specimens), 3.9 (3.8-4.0) grams; basilar length 43.8 (41.9-47.0); postorbital breadth more than width of basioccipital measured from medial margin of one foramen lacerum posterior to its opposite; interorbital breadth not less than distance between foramen opticum and anterior margin of tympanic bulla; anterior margin of tympanic bulla as far posterior to foramen ovale as width of 2 to 2-1/2 (including I3) upper incisors; length of tympanic bulla more than length of lower molar and premolar tooth-row and longer than rostrum; anterior margin of masseteric fossa below m2.Female (based on 4 adults from San Diego County): See measurements. As described inM. f. nigriaurisexcept that: Weight, 2.6 (2.2-2.8) grams; basilar length, 40.0 and 40.1; postorbital breadth more than width of basioccipital measured from medial margin of one foramen lacerum posterior to its opposite; length of tympanic bulla more than length of rostrum.The skull of the female averages 34 per cent lighter than that of the average male.
Type.—Male, adult, skull and skin; no. 3257, Mus. Vert. Zoöl.; San Diego, San Diego County, California; May 20, 1907; obtained by Frank X. Holzner.
Right M1 is missing and the part of the jaw bearing this tooth is broken away. With this exception the skull is complete and unbroken and the teeth are all present and entire. The skin is fairly well made and in good condition except that it is slightly soiled.
Range.—Altitudinally sea level to 8000 feet (Tahquitz Valley, San Jacinto Mountains); Sonoran and Transition life-zones of coast and mountains west of Mohave and Imperial deserts of southern California from Point Conception and Cuyama Valley southward at least to Mexican boundary. See figures29and30on pages 221 and 314.
Characters for ready recognition.—Differs fromM. f. nigriaurisby having postorbital breadth of adult males and females, more, rather than less, than width of basioccipital measured from medial margin of one foramen lacerum posterior to its opposite; fromM. f. pulchraby having tympanic bulla longer than rostrum (orbitonasal length) and by near (l) Antique Brown rather than near (16j) Buckthorn Brown to near (h) Yellow Ocher color of upper parts.
Description.—Size.—Male: Six adults and subadults from San Diego yield average and extreme measurements as follows: Total length, 439 (428-449); length of tail, 153 (142-160); length of hind foot, 45 (40-47). Corresponding measurements for a series of eight adult males from the vicinity of Los Angeles are: 416 (394-428); 158 (151-166); 44 (40-47). In the series from San Diego the tail averages 54 per cent as long as head and body. In the series from Los Angeles the average is 61 per cent. Length of hind foot in each series, less than basal length. The type specimen measures, 435, 142, 42.
Female: No. 5070, adult, from San Diego, measures 367, 141, 38. Nos. 22 and 6748 from Santa Ysabel, measure: 359, 380; 130, 140; 39, 35. No. 7194 from Jamacha measures, 358, 125, 35. Three adult females from Los Angeles yield the following: Total length, 373, 345, 368; length of tail, 150, 122, 134; length of hind foot,—, 41, 41. In no. 5070 the tail is 62 per cent as long as the head and body and in the three from Los Angeles it averages 60 (55-67) per cent. Length of hind foot, in each case, less than basal length.
The average differences in external measurements of the two sexes as shown by the six males from San Diego and the four females from San Diego County are: Total length, 73; length of tail, 19; length of hind foot, 8. Corresponding differences shown by the eight males and three females from Los Angeles are: 54, 23, 3.
Externals.—Longest facial vibrissae brownish, like dark color of head and extending beyond ear; carpal vibrissae mostly color of underparts and extending to apical pad of fifth digit; hairiness of foot-soles slightly more than shown in figure20.
Color.—Spot between eyes, band confluent with color of underparts on each side of head extending anterodorsally anterior to ear, and posterior third of each upper lip tinged with color of underparts or, less often, white; chin and lower lips white; remainder of sides and top of head posteriorly to near line connecting posterior margins of ears, blackish; inside of pinna of ear, and sometimes outside of pinna, blackish; dark spot posterior to each angle of mouth present on each side in three-fourths of specimens; tip of tail black; remainder of upper parts near (l) Antique Brown, and with more yellow than tone 3 of Raw Umber of Oberthür and Dauthenay, pl. 301. Underparts Ochraceous-Buff to Warm Buff and in some specimens from Los Angeles and Ventura counties Ochraceous-Orange, especially in young and juveniles. Color of underparts extends distally on posterior sides of forelegs over toes onto antipalmar faces of feet and wrists and on medial sides of hind limbs over antiplantar faces of toes. Least width of color of underparts averaging, in 15 adult and subadult males from San Diego County, 54 (35-75) per cent of greatest width of color of upper parts. Black tip of tail in same series of males averaging 54.5 (46-60) mm. long. Thus averaging longer than hind foot and 35 per cent of length of tail-vertebrae.
Skull and teeth.—Male (based on 6 adults from San Diego County). See measurements and plates22-24. As described inM. f. nigriaurisexcept that: Weight (4 specimens), 3.9 (3.8-4.0) grams; basilar length 43.8 (41.9-47.0); postorbital breadth more than width of basioccipital measured from medial margin of one foramen lacerum posterior to its opposite; interorbital breadth not less than distance between foramen opticum and anterior margin of tympanic bulla; anterior margin of tympanic bulla as far posterior to foramen ovale as width of 2 to 2-1/2 (including I3) upper incisors; length of tympanic bulla more than length of lower molar and premolar tooth-row and longer than rostrum; anterior margin of masseteric fossa below m2.
Female (based on 4 adults from San Diego County): See measurements. As described inM. f. nigriaurisexcept that: Weight, 2.6 (2.2-2.8) grams; basilar length, 40.0 and 40.1; postorbital breadth more than width of basioccipital measured from medial margin of one foramen lacerum posterior to its opposite; length of tympanic bulla more than length of rostrum.
The skull of the female averages 34 per cent lighter than that of the average male.
The skull of the male oflatirostra, compared with that ofnigriauris, is by weight, more than one-fourth lighter, has a lesser basilar length, a lesser mastoid breadth, a lesser zygomatic breadth and a narrower M1. In these features no overlap has been observed between adults from the general vicinities of the type localities of the two forms. In adult males oflatirostrathe postorbital breadth, with one exception, is more than the combined length of P4 and P3 whereas the reverse is true in adult males ofnigriauris. Both males and females oflatirostrahave a generally smaller skull with relatively broader interorbital and postorbital parts and the tympanic bullae are relatively larger, rounder and more inflated.
Compared with the skull of the male ofpulchrathat oflatirostrais, by weight, more than one-fourth lighter, has a lesser basilar and orbitonasal length, lesser zygomatic and mastoid breadth and a more nearly flat braincase. In these features no overlap has been observed between adults from the general vicinities of the type localities of the two subspecies. Also, inlatirostrathe tympanic bulla is longer than the rostrum whereas the opposite is true inpulchra. The skull oflatirostrais generally smaller and relatively, on the average, has the preorbital part of the skull deeper and broader with longer tooth-rows, although with shorter rostrum, while the zygomatic and mastoid breadths are less. Study of skulls of subadult females ofpulchraindicate that females oflatirostraandpulchradiffer in the same fashion as do males.
Remarks.—This subspecies long has gone by the nameM. xanthogenysand the type locality was generally supposed to be in the vicinity of San Diego. This supposition seems to have originated with Merriam's (1896:25) statement that the type locality was "Southern California, probably vicinity of San Diego." Nevertheless, as set forth in the account ofM. f. xanthogenysthe type specimen concerned now is thought to have come from much farther north.
Although 76 Recent specimens are available from southern California, additional adults are needed to understand the geographic variation there.M. f. latirostramay be a composite—made up of more than one geographic race. Specimens from San Diego County differ so much in relative length of the tail that at one stage in the present study it was thought that a difference in this respect existed between the coastal animals and those from farther inland. Material received later did not wholly substantiate this view and because of the uniformly small size of all of the skulls from that county, the animals were later regarded as of the same subspecies. Eventually, even this supposed common feature proved to be inconstant for an adult male from Jamacha, no. 7098, of the San Diego Society of Natural History, and another adult male from San Marcos, no. 8869, collection of Ralph Ellis, were later examined and found to have skulls as large as those of average-sized, adult males ofnigriauris.
Despite these puzzling local variations, it is established that the long-tailed weasels of southern California are smaller than those from farther north. Also, the southern animal averages smaller in weight and size of skull, and the skull is differently proportioned. Specimens in series from Los Angeles County definitely are intermediate in size and shape of skull betweenlatirostrafrom San Diego County andnigriaurisfrom, say, Santa Clara County, but definitely more closely resemblelatirostrafrom San Diego County than they donigriauris. A skull of a young animal, not here identified to subspecies, from Potholes, in the Colorado River Valley, 10 miles northeast of Bard, Imperial County, California, may have closest relationship toM. f. latirostra. Additional comment on this specimen is offered in the account ofM. f. neomexicana.
From the asphalt pits of Rancho La Brea, in Los Angeles County, a total of 57 skulls have been examined, more than half of which are reasonably complete. I have been unable to learn whether these came from pits regarded by students of the deposit as wholly Recent, from pits regarded as of Pleistocene age, or from both. Suffice to say that only two specimens were found which could be distinguished from skulls of the subspecies of weasel living in that area today.
These two specimens, lent to me by Professor Chester Stock, were with other skulls received from the Los Angeles Museum of History, Art and Science and bore identifying numbers as follows: 16/20-27, the anterior part of the skull of an adult, and 16, the skull posterior to the cribiform plate of a subadult or possibly young individual. The latter has a mastoid breadth of 28.0 millimeters, a tympanic bulla 16.1 long and other measurements in proportion. It is larger than any specimen of weasel, of any subspecies, seen from California and in the subgenusMustelaseems to be exceeded in size only by certain individuals ofM. f. texensis.M. f. neomexicanaattains relatively large size and comparisons were made with individuals of that subspecies. However, the young specimen from Rancho La Brea differs fromneomexicanain that the tympanic bullae rise less steeply on the medial sides and the inferior lip of the external auditory meatus is less developed laterally. Age considered, the sagittal crest is less developed and the mastoid processes project more abruptly from the skull. The anterior part of the skull of the adult, no. 16/20-27 is larger than any specimen seen ofM. f. latirostraor adjoining subspecies, and among California-taken specimens is equaled in size only by the largest males ofM. f. mundafrom the northwest coastal district in Mendocino County. This adult from Rancho La Brea differs fromneomexicana, sex and age taken into account, in greater postorbital breadth, lesser rostral width in comparison with the interorbital breadth, and in having the temporal ridges at the anterior end of the sagittal crest spread out into a Y-shaped, rather than a T-shaped, pattern. All these differences fromneomexicanaare features of agreement with the California bridled weasels of the subspecieslatirostra,nigriauris, andmunda. The same is true of the characters which set apart the young specimen fromneomexicana. In summary: of 57 weasel skulls examined from the asphalt pits at Rancho La Brea, Los Angeles County, all but two are indistinguishable from the skulls of the Recent weasel living in that region today. These two skulls agree in qualitative characters with animals of the California coastal subspecies now living from Los Angeles northward to Humboldt County, but are larger. For the time being these two may be thought of as giants of the same type of animal inhabiting the Los Angeles region today.
Only one of 41 adult and subadult skulls examined for malformation of the frontal sinuses shows infestation by parasites.
Specimens examined.—Total number, 142, listed by counties from north to south. Unless otherwise indicated specimens are in the Museum of Vertebrate Zoölogy.California.Santa Barbara County: Rincon Point, 1.Ventura County: Cuyama Valley, 2200 ft., 1[91]; Nordhoff, 3[59]; Santa Paula, 1[59]; Ventura, 7.Los Angeles County: near Owensmouth, 1[24]; Cahuenga, 1[91]; Llano, 10 mi. E Littlerock, 1; Flint Ridge, Pasadena, 1[59]; Pasadena, 3; Lankershim, 1[24]; 1 mi. S Lankershim, 1[24]; Duarte, 1[59]; Covina, 1[59]; Claremont, 1[91]; El Monte, 4 (2[75], 1[24]); Montebello, 1; Alhambra, 6 (5[2], 1[91]); El Nogal, 2[8]; Gardena, 1[26]; Palos Verdes Estate, 3; Rancho La Brea asphalt deposits, 57[70]and[92].San Bernardino County: San Bernardino Valley, 1[75]; San Bernardino, 4 (2[20], 1[91]); Redlands, 2 (1[38]); Bluff Lake, 2 (1[59], 1[33]).Riverside County: West Riverside, 1; Arlington, 800 ft., 1[17]; 3-1/2 mi. E and 1/2 mi. N Beaumont, 2600 ft., 1; Banning, 1[91]; Cabazon, 1[91]; San Jacinto Plain, 1[20]; Tahquitz Valley, 8000 ft., 1; Elsinore, 1[1].San Diego County: Twin Oaks, 1[91]; San Marcos, 2 (1[87], 1[41]); Escondido, 1; Witch Creek, 1[91]; Ballena, 1[20]; Santa Ysabel, 3 (2[20], 1[87]); Julián, 1; La Jolla, 1; Lakeside, 1[91]; El Cajon, 1[91]; El Vido (not found on map), 1[91]; San Diego, 9 (1[91], 1[20], 1[87], 1[32]); Jamacha, 2[87]; Chula Vista, 1[20].
Specimens examined.—Total number, 142, listed by counties from north to south. Unless otherwise indicated specimens are in the Museum of Vertebrate Zoölogy.
California.Santa Barbara County: Rincon Point, 1.Ventura County: Cuyama Valley, 2200 ft., 1[91]; Nordhoff, 3[59]; Santa Paula, 1[59]; Ventura, 7.Los Angeles County: near Owensmouth, 1[24]; Cahuenga, 1[91]; Llano, 10 mi. E Littlerock, 1; Flint Ridge, Pasadena, 1[59]; Pasadena, 3; Lankershim, 1[24]; 1 mi. S Lankershim, 1[24]; Duarte, 1[59]; Covina, 1[59]; Claremont, 1[91]; El Monte, 4 (2[75], 1[24]); Montebello, 1; Alhambra, 6 (5[2], 1[91]); El Nogal, 2[8]; Gardena, 1[26]; Palos Verdes Estate, 3; Rancho La Brea asphalt deposits, 57[70]and[92].San Bernardino County: San Bernardino Valley, 1[75]; San Bernardino, 4 (2[20], 1[91]); Redlands, 2 (1[38]); Bluff Lake, 2 (1[59], 1[33]).Riverside County: West Riverside, 1; Arlington, 800 ft., 1[17]; 3-1/2 mi. E and 1/2 mi. N Beaumont, 2600 ft., 1; Banning, 1[91]; Cabazon, 1[91]; San Jacinto Plain, 1[20]; Tahquitz Valley, 8000 ft., 1; Elsinore, 1[1].San Diego County: Twin Oaks, 1[91]; San Marcos, 2 (1[87], 1[41]); Escondido, 1; Witch Creek, 1[91]; Ballena, 1[20]; Santa Ysabel, 3 (2[20], 1[87]); Julián, 1; La Jolla, 1; Lakeside, 1[91]; El Cajon, 1[91]; El Vido (not found on map), 1[91]; San Diego, 9 (1[91], 1[20], 1[87], 1[32]); Jamacha, 2[87]; Chula Vista, 1[20].
Long-tailed Weasel
Plates22,23and24
Mustela frenata pulchraHall, Carnegie Instit. Washington Publ. 473:98, November 20, 1936.
Mustela frenata pulchraHall, Carnegie Instit. Washington Publ. 473:98, November 20, 1936.
Type.—Male, adult, skeleton and skin; no. 16668, Mus. Vert. Zoöl.; Buttonwillow, Kern County, California; April 30, 1912; obtained by J. Grinnell; original no. 1953.The skull (plates22-24) is complete and unbroken (a fracture in the right jugal has healed). All teeth are present and entire. The skeleton lacks the os penis, left fibula, shaft of left tibia and the distal three or four caudal vertebrae. Some of the bones of the feet distal to the radius and tibia are with the skeleton, and the remainder probably are in the skin. The skin is fairly well made and in good condition, except for the left hind leg which was torn when the animal was captured. A well-developed scrotal pouch shows the specimen to have been a male.Range.—Altitudinally around 300 feet in San Joaquin Valley to 2500 feet at Isabella; Upper Sonoran and Lower Sonoran life-zones of southern end of San Joaquin Valley and in mountains at southern end of Valley, California. See figures29and30on pages 221 and 314.Characters for ready recognition.—Differs fromM. f. nevadensisin presence of light facial markings, and fromM. f. nevadensisandM. f. inyoensisin near (16j) Buckthorn Brown to near (h) Yellow Ocher rather than near (14ntol) Brussels Brown color of upper parts, and greater size with hind foot more than 40 in females and basilar length averaging more than 46.0 in males; fromM. f. latirostrain having rostrum (orbitonasal length) longer than tympanic bulla and fromM. f. latirostraandM. f. nigriaurisby color of upper parts as stated above rather than near (l) Antique Brown, and by having inside of ears same color as back rather than much darker than back; fromM. f. xanthogenysin hind foot of males more than 46 and broader skull which in males has breadth of rostrum more than 13.9 and mastoid breadth more than 26.0.Description.—Size.—Male: The type specimen and five other adults yield average and extreme measurements as follows: Total length, 454 (428-477); length of tail, 178 (153-184); length of hind foot, 50 (47-55). Tail averages 65 per cent as long as head and body. Length of hind foot approximately equal to basal length. The type specimen measures, 460, 184, 49.Female: Three subadult topotypes yield average and extreme measurements as follows: Total length, 399 (383-411); length of tail, 154 (140-161); length of hind foot, 42 (41-42). Tail averages 63 per cent as long as head and body. Length of hind foot less than basal length.The average differences in external measurements of the two sexes are: Total length, 55; length of tail, 24; length of hind foot, 8.Externals.—As described inMustela frenata nigriauris.Color.—Spot between eyes, band confluent with color of underparts, on each side of head extending anterodorsally anterior to each ear, posterior third of each upper lip, lower lips and chin white or more often darker than Ochraceous-Buff and therefore same color as belly; dark spot posterior to each angle of mouth present but small; tip of tail black; remainder of upper parts near (16j) Buckthorn Brown to near (h) Yellow Ocher and from tone 2 to 4 of Brown Pink of Oberthür and Dauthenay, pl. 297, but with a trifle more reddish brown. Upper parts of uniform color except for occasional slight darkening of nose, forehead, and areas around eyes. Underparts darker (a) than Ochraceous-Buff. Color of underparts extends distally on posterior sides of forelegs over toes, onto antipalmar faces of feet and wrists, on medial sides of hind limbs over antiplantar faces of toes, tarsal region and sometimes in diluted fashion on proximal third of underside of tail. Least width of color of underparts averaging, in 6 male topotypes, 55 (43-81) per cent of greatest width of color of upper parts. Black tip of tail in same series of males averaging 58 (53-63) mm. long; thus averaging longer than hind foot and 33 per cent of length of tail-vertebrae.Skull and teeth.—Male (based on 6 ads., type and 5 topotypes): See measurements and plates22-24. As described inM. f. nigriaurisexcept that: Weight (6 ads.), 5.3 (4.5-6.1) grams; basilar length, 47.6 (46.0-48.6); (one skull, no. 335, with postorbital breadth more than distance between posterior borders of P4 and P2); interorbital breadth more or less than distance between foramen opticum and anterior margin of tympanic bulla; anterior margin of tympanic bulla as far posterior to foramen ovale as width of 2 to 3-1/2 (including I3) upper incisors; length of tympanic bulla more than length of lower molar and premolar tooth-row and shorter than rostrum.Female: Adult skull of typical female not seen.
Type.—Male, adult, skeleton and skin; no. 16668, Mus. Vert. Zoöl.; Buttonwillow, Kern County, California; April 30, 1912; obtained by J. Grinnell; original no. 1953.
The skull (plates22-24) is complete and unbroken (a fracture in the right jugal has healed). All teeth are present and entire. The skeleton lacks the os penis, left fibula, shaft of left tibia and the distal three or four caudal vertebrae. Some of the bones of the feet distal to the radius and tibia are with the skeleton, and the remainder probably are in the skin. The skin is fairly well made and in good condition, except for the left hind leg which was torn when the animal was captured. A well-developed scrotal pouch shows the specimen to have been a male.
Range.—Altitudinally around 300 feet in San Joaquin Valley to 2500 feet at Isabella; Upper Sonoran and Lower Sonoran life-zones of southern end of San Joaquin Valley and in mountains at southern end of Valley, California. See figures29and30on pages 221 and 314.
Characters for ready recognition.—Differs fromM. f. nevadensisin presence of light facial markings, and fromM. f. nevadensisandM. f. inyoensisin near (16j) Buckthorn Brown to near (h) Yellow Ocher rather than near (14ntol) Brussels Brown color of upper parts, and greater size with hind foot more than 40 in females and basilar length averaging more than 46.0 in males; fromM. f. latirostrain having rostrum (orbitonasal length) longer than tympanic bulla and fromM. f. latirostraandM. f. nigriaurisby color of upper parts as stated above rather than near (l) Antique Brown, and by having inside of ears same color as back rather than much darker than back; fromM. f. xanthogenysin hind foot of males more than 46 and broader skull which in males has breadth of rostrum more than 13.9 and mastoid breadth more than 26.0.
Description.—Size.—Male: The type specimen and five other adults yield average and extreme measurements as follows: Total length, 454 (428-477); length of tail, 178 (153-184); length of hind foot, 50 (47-55). Tail averages 65 per cent as long as head and body. Length of hind foot approximately equal to basal length. The type specimen measures, 460, 184, 49.
Female: Three subadult topotypes yield average and extreme measurements as follows: Total length, 399 (383-411); length of tail, 154 (140-161); length of hind foot, 42 (41-42). Tail averages 63 per cent as long as head and body. Length of hind foot less than basal length.
The average differences in external measurements of the two sexes are: Total length, 55; length of tail, 24; length of hind foot, 8.
Externals.—As described inMustela frenata nigriauris.
Color.—Spot between eyes, band confluent with color of underparts, on each side of head extending anterodorsally anterior to each ear, posterior third of each upper lip, lower lips and chin white or more often darker than Ochraceous-Buff and therefore same color as belly; dark spot posterior to each angle of mouth present but small; tip of tail black; remainder of upper parts near (16j) Buckthorn Brown to near (h) Yellow Ocher and from tone 2 to 4 of Brown Pink of Oberthür and Dauthenay, pl. 297, but with a trifle more reddish brown. Upper parts of uniform color except for occasional slight darkening of nose, forehead, and areas around eyes. Underparts darker (a) than Ochraceous-Buff. Color of underparts extends distally on posterior sides of forelegs over toes, onto antipalmar faces of feet and wrists, on medial sides of hind limbs over antiplantar faces of toes, tarsal region and sometimes in diluted fashion on proximal third of underside of tail. Least width of color of underparts averaging, in 6 male topotypes, 55 (43-81) per cent of greatest width of color of upper parts. Black tip of tail in same series of males averaging 58 (53-63) mm. long; thus averaging longer than hind foot and 33 per cent of length of tail-vertebrae.
Skull and teeth.—Male (based on 6 ads., type and 5 topotypes): See measurements and plates22-24. As described inM. f. nigriaurisexcept that: Weight (6 ads.), 5.3 (4.5-6.1) grams; basilar length, 47.6 (46.0-48.6); (one skull, no. 335, with postorbital breadth more than distance between posterior borders of P4 and P2); interorbital breadth more or less than distance between foramen opticum and anterior margin of tympanic bulla; anterior margin of tympanic bulla as far posterior to foramen ovale as width of 2 to 3-1/2 (including I3) upper incisors; length of tympanic bulla more than length of lower molar and premolar tooth-row and shorter than rostrum.
Female: Adult skull of typical female not seen.
As compared with the skull of the type specimen ofinyoensis, skulls of adult males ofpulchraare larger throughout, relatively broader, especially in the preorbital part of the skull, have more inflated tympanic bullae, and are less convex in dorsal outline. Comparison of the skull with that oflatirostrahas been made in discussion of that subspecies. Comparison of skulls of adult males ofnigriaurisandpulchrashows that those ofpulchraaverage larger in every measurement taken except those of m1, M1, P4, and depth of skull at posterior borders of upper molars. The basilar length is only slightly more and it follows that, relative to this length, other measurements of the skull are relatively, as well as actually, larger. In no one measurement is there an entire lack of overlap, but the skulls of adult males, and probably adult females, may be distinguished from those ofnigriaurisby the combination of the following mentioned, average differences: Tympanic bullae larger in each of three dimensions; preorbital and interorbital parts of skull broader and notably heavier; interorbital breadth greater; zygomatic arches more expanded laterally; mastoid processes more prominent. As compared withxanthogenys, differences of similar nature, but of greater degree, distinguishpulchra. As compared with those ofnevadensis(represented by specimens from Mono Co., Calif.), skulls of adult males ofpulchraaverage larger in every measurement taken and no overlap exists in basilar length, orbitonasal length, mastoid breadth, zygomatic breadth, length of tympanic bulla, or depth of skull at either the anterior margin of the basioccipital or at the posterior margins of the upper molars. Relatively, the preorbital portion is about the same size in the two forms.
Remarks.—The best material of this big weasel was obtained in 1910 and 1911 by John Wimmer and forwarded to the California Academy of Sciences through John R. Rowley, although in 1905, one specimen had been obtained by A. S. Bunnell for the collections of the United States Bureau of Biological Survey, another by J. Grinnell for the Museum of Vertebrate Zoölogy in 1912, and in 1933, another by L. M. Huey, for the San Diego Society of Natural History.
The males from the type locality are relatively uniform in size and shape of skull. The one exception is no. 137935, U. S. Nat. Mus., slightly younger than the others. Its skull is relatively more slender than any of the others and does not display several of the differential characters. The male, no. 127566, U. S. Nat. Mus., from Alila (= Earlimart) is intermediate in cranial features betweenpulchraandxanthogenysas known from specimens taken in the vicinity of Fresno. The skull of the female, no. 127565, from the same place, is too young to provide diagnostic characters. Since the skull of an adult female of topotypicalpulchrais unknown, doubt attaches to the identification of the adult, female specimen, no. 115895, U. S. Nat. Mus., from Delano. It has a relatively broad skull in comparison with the adult female ofxanthogenysfrom Los Banos. The adult female, no. 9998, San Diego Soc. Nat. Hist., from 2 mi. SW Simmler, shows approach tonigriaurisin slightly reduced size. The skin alone from Coalinga, a male, taken on April 10, 1935, measures 462, 179, 47. The adult female, with crushed skull, from 4 miles east of Coalinga, measures 350, 129, 40. In size, these specimens agree better withpulchrathan withxanthogenys. The skin alone from 3 miles south of Coalinga is unsexed and without external measurements. Skulls of adults from Coalinga are needed to permit of more positive identification of the subspecies found there. The female from 4 miles east of Coalinga, taken on February 21, 1936, is in process of molt on the underparts, and the longer hair which is near (20´) Naples Yellow contrasts strongly with the incoming shorter hair which is near (10c) Salmon-Orange. The skin alone, no. 16270, Mus. Vert. Zoöl., from Isabella, was made up from a decayed animal and is of but little use. It is referred topulchrapurely because of geographic nearness of Isabella to the type locality ofpulchra. The most that can be told from the specimen is that it is a relatively light-colored, bridled weasel. The fact that the color is slightly darker than inpulchramay or may not indicate intergradation withnevadensis. No. 54103/41042, U. S. Nat. Mus., consisting of crushed bits of skull and the skin of the head, is from Willow Spring, Kern County. This marginal locality is really in the Mojave Desert rather than in the San Joaquin Valley. The light color of the skin of the head suggestspulchra, but it is realized that a complete specimen might show the animal there to be unlikepulchra.
None of the skulls shows evidence of having had the frontal sinuses infested by parasites.
Specimens examined.—Total number, 18, listed by counties from north to south. Unless otherwise indicated, specimens are in the Museum of Vertebrate Zoölogy.California.Fresno County: Coalinga, 1[23]; 4 mi. E Coalinga, 1; 3 mi. S Coalinga, 1[8].Tulare County: Alila (= Earlimart), 2[91].Kern County: Delano, 1[91]; Buttonwillow, 9 (6[8], 2[91]); Isabella, 1; Willow Spring, 1[91]San Luis Obispo County: 2 mi. SW Simmler, 1[87].
Specimens examined.—Total number, 18, listed by counties from north to south. Unless otherwise indicated, specimens are in the Museum of Vertebrate Zoölogy.
California.Fresno County: Coalinga, 1[23]; 4 mi. E Coalinga, 1; 3 mi. S Coalinga, 1[8].Tulare County: Alila (= Earlimart), 2[91].Kern County: Delano, 1[91]; Buttonwillow, 9 (6[8], 2[91]); Isabella, 1; Willow Spring, 1[91]San Luis Obispo County: 2 mi. SW Simmler, 1[87].
Long-tailed Weasel
Plates22,23and24
Mustela frenata inyoensisHall, Carnegie Instit. Washington Publ. 473:99, November 20, 1936.Putorius xanthogenys, Merriam, N. Amer. Fauna, 11:25, June 30, 1896 (part).Mustela xanthogenys xanthogenys, Miller, U. S. Nat. Mus. Bull., 79:99, December 31, 1912.
Mustela frenata inyoensisHall, Carnegie Instit. Washington Publ. 473:99, November 20, 1936.
Putorius xanthogenys, Merriam, N. Amer. Fauna, 11:25, June 30, 1896 (part).
Mustela xanthogenys xanthogenys, Miller, U. S. Nat. Mus. Bull., 79:99, December 31, 1912.
Type.—Male, adult, skull (with skeleton) and skin; no. 25907, Mus. Vert. Zoöl.; Carl Walter's Ranch, 2 mi. N Independence, Inyo County, California; June 26, 1917; obtained by A. C. Shelton; original no. 3143.The skull (plates22-24) is complete and unbroken. All teeth are present and entire. The skin is well made and in good condition.Range.—From 3700 feet (Lone Pine) to at least 4000 feet (Alvord); Lower Sonoran Life-zone of the floor of Owens Valley in Inyo County, California. See figures29and30on pages 221 and 314.Characters for ready recognition.—Differs fromM. f. nevadensisin presence of white facial markings; fromM. f. pulchrain near (l) Brussels Brown rather than near (16j) Buckthorn Brown to near (h) Yellow Ocher color of upper parts and basilar length of less than 45 in males; fromM. f. latirostrain brownish rather than blackish color of inside of ear and orbitonasal length of more than 15.Description.—Size.—Male: Two adults, the type specimen and no. 25392/32805, measure, respectively, as follows: Total length, 423 and 390; length of tail, 170 and 145; length of hind foot, 42 and 44. Tail is 67 and 59 per cent as long as head and body. Length of hind foot less than basal length.Female: No. 12400, Field Mus. Nat. Hist., which is young, has the following measurements: Total length, 390; length of tail, 150; length of hind foot, 39. Tail is 63 per cent as long as head and body. Length of hind foot less than basal length.The differences in external measurements between the two sexes, as represented by the male type specimen and by the young female, are: Total length, 33, length of tail, 20; length of hind foot, 3.Externals.—Longest facial vibrissae black or dark brown and reaching beyond ear; carpal vibrissae same color as underparts and extending to apical pad of fifth digit; hairiness of foot-soles (in summer pelage) slightly less than shown in figure19.Color.—Large spot between eyes, band confluent with color of underparts, on each side of head extending anterodorsally anterior to each ear, upper throat, chin, lower lips and in some specimens part or all of upper lips white; patch between eyes and bars in front of ears tinged with some shade of yellowish in one specimen; dark spot posterior to each angle of mouth present in four of five specimens; tip of tail black; remainder of upper parts, in summer, near (l) Brussels Brown or tones 1 to 2 of Raw Umber of Oberthür and Dauthenay, pl. 301; slightly darker brown on forehead, nose and about eyes. In winter near (j) Snuff Brown or lighter than Brussels Brown with a smoked effect. Underparts Buff-Yellow, winter and summer. Color of underparts extends distally on posterior sides of forelegs over toes onto antipalmar faces of feet and wrists and on medial sides of hind legs over antiplantar faces of toes. Least width of color of underparts averaging, in 5 available specimens 34 (24-42) per cent of greatest width of color of upper parts. Black tip of tail, in two adult males, averaging 53 (45 and 60) mm. Thus longer than hind foot and averaging 34 per cent of length of tail-vertebrae.Skull and teeth.—Male (based on the type): See measurements and plates22-24. As described inM. f. nigriaurisexcept that: Weight, 4.4 grams; basilar length, 44.7; postorbital breadth not less than width of basioccipital measured from medial margin of one foramen lacerum posterior to its opposite; length of tympanic bulla less than length of lower molar and premolar tooth-row.Female: Adult unknown.
Type.—Male, adult, skull (with skeleton) and skin; no. 25907, Mus. Vert. Zoöl.; Carl Walter's Ranch, 2 mi. N Independence, Inyo County, California; June 26, 1917; obtained by A. C. Shelton; original no. 3143.
The skull (plates22-24) is complete and unbroken. All teeth are present and entire. The skin is well made and in good condition.
Range.—From 3700 feet (Lone Pine) to at least 4000 feet (Alvord); Lower Sonoran Life-zone of the floor of Owens Valley in Inyo County, California. See figures29and30on pages 221 and 314.
Characters for ready recognition.—Differs fromM. f. nevadensisin presence of white facial markings; fromM. f. pulchrain near (l) Brussels Brown rather than near (16j) Buckthorn Brown to near (h) Yellow Ocher color of upper parts and basilar length of less than 45 in males; fromM. f. latirostrain brownish rather than blackish color of inside of ear and orbitonasal length of more than 15.
Description.—Size.—Male: Two adults, the type specimen and no. 25392/32805, measure, respectively, as follows: Total length, 423 and 390; length of tail, 170 and 145; length of hind foot, 42 and 44. Tail is 67 and 59 per cent as long as head and body. Length of hind foot less than basal length.
Female: No. 12400, Field Mus. Nat. Hist., which is young, has the following measurements: Total length, 390; length of tail, 150; length of hind foot, 39. Tail is 63 per cent as long as head and body. Length of hind foot less than basal length.
The differences in external measurements between the two sexes, as represented by the male type specimen and by the young female, are: Total length, 33, length of tail, 20; length of hind foot, 3.
Externals.—Longest facial vibrissae black or dark brown and reaching beyond ear; carpal vibrissae same color as underparts and extending to apical pad of fifth digit; hairiness of foot-soles (in summer pelage) slightly less than shown in figure19.
Color.—Large spot between eyes, band confluent with color of underparts, on each side of head extending anterodorsally anterior to each ear, upper throat, chin, lower lips and in some specimens part or all of upper lips white; patch between eyes and bars in front of ears tinged with some shade of yellowish in one specimen; dark spot posterior to each angle of mouth present in four of five specimens; tip of tail black; remainder of upper parts, in summer, near (l) Brussels Brown or tones 1 to 2 of Raw Umber of Oberthür and Dauthenay, pl. 301; slightly darker brown on forehead, nose and about eyes. In winter near (j) Snuff Brown or lighter than Brussels Brown with a smoked effect. Underparts Buff-Yellow, winter and summer. Color of underparts extends distally on posterior sides of forelegs over toes onto antipalmar faces of feet and wrists and on medial sides of hind legs over antiplantar faces of toes. Least width of color of underparts averaging, in 5 available specimens 34 (24-42) per cent of greatest width of color of upper parts. Black tip of tail, in two adult males, averaging 53 (45 and 60) mm. Thus longer than hind foot and averaging 34 per cent of length of tail-vertebrae.
Skull and teeth.—Male (based on the type): See measurements and plates22-24. As described inM. f. nigriaurisexcept that: Weight, 4.4 grams; basilar length, 44.7; postorbital breadth not less than width of basioccipital measured from medial margin of one foramen lacerum posterior to its opposite; length of tympanic bulla less than length of lower molar and premolar tooth-row.
Female: Adult unknown.