Chapter 18

LEOPOLD I.(1790-1865), king of the Belgians, fourth son of Francis, duke of Saxe-Coburg-Saalfeld, and uncle of Queen Victoria of England, was born at Coburg on the 18th of December 1790. At the age of eighteen he entered the military service of Russia, and accompanied the emperor Alexander to Erfurt as a member of his staff. He was required by Napoleon to quit the Russian army, and spent some years in travelling. In 1813 he accepted from the emperor Alexander the post of a cavalry general in the army of invasion, and he took part in the whole of the campaign of that and the following year, distinguishing himself in the battles of Leipzig, Lützen and Bautzen. He entered Paris with the allied sovereigns, and accompanied them to England. He married in May 1816 Charlotte, only child of George, prince regent, afterwards George IV., heiress-presumptive to the British throne, and was created duke of Kendal in the British peerage and given an annuity of £50,000. The death of the princess in the following year was a heavy blow to his hopes, but he continued to reside in England. In 1830 he declined the offer of the crown of Greece, owing to the refusal of the powers to grant conditions which he considered essential to the welfare of the new kingdom, but was in the following year elected king of the Belgians (4th June 1831). After some hesitation he accepted the crown, having previously ascertained that he would have the support of the great powers on entering upon his difficult task, and on the 12th of July he made his entry into Brussels and took the oath to observe the constitution. During the first eight years of his reign he was confronted with the resolute hostility of King William I. of Holland, and it was not until 1839 that the differences between the two states, which until 1830 had formed the kingdom of the Netherlands, were finally settled at the conference of London by the treatyof the 24 Articles (seeBelgium). From this date until his death, King Leopold spent all his energies in the wise administration of the affairs of the newly formed kingdom, which may be said to owe in a large measure its first consolidation and constant prosperity to the care and skill of his discreet and fatherly government. In 1848 the throne of Belgium stood unshaken amidst the revolutions which marked that year in almost every European country. On the 8th of August 1832 Leopold married, as his second wife, Louise of Orleans, daughter of Louis Philippe, king of the French. Queen Louise endeared herself to the Belgian people, and her death in 1850 was felt as a national loss. This union produced two sons and one daughter—(1) Leopold, afterwards king of the Belgians; (2) Philip, count of Flanders; (3) Marie Charlotte, who married Maximilian of Austria, the unfortunate emperor of Mexico. Leopold I. died at Laeken on the 10th of December 1865. He was a most cultured man and a great reader, and did his utmost during his reign to encourage art, science and education. His judgment was universally respected by contemporary sovereigns and statesmen, and he was frequently spoken of as “the Nestor of Europe” (see alsoVictoria, Queen).

See Th. Juste,Léopold Ier, roi des Belges d’après des doc. inéd. 1793-1865(2 vols., Brussels, 1868), andLes Fondateurs de la monarchie Belge(22 vols., Brussels, 1878-1880); J. J. Thonissen,La Belgique sous le règne de Léopold Ier(Louvain, 1862).

See Th. Juste,Léopold Ier, roi des Belges d’après des doc. inéd. 1793-1865(2 vols., Brussels, 1868), andLes Fondateurs de la monarchie Belge(22 vols., Brussels, 1878-1880); J. J. Thonissen,La Belgique sous le règne de Léopold Ier(Louvain, 1862).

LEOPOLD II.[Leopold Louis Philippe Marie Victor] (1835-1909), king of the Belgians, son of the preceding, was born at Brussels on the 9th of April 1835. In 1846 he was created duke of Brabant and appointed a sub-lieutenant in the army, in which he served until his accession, by which time he had reached the rank of lieutenant-general. On attaining his majority he was made a member of the senate, in whose proceedings he took a lively interest, especially in matters concerning the development of Belgium and its trade. On the 22nd of August 1853 Leopold married Marie Henriette (1836-1902), daughter of the archduke Joseph of Austria, palatine of Hungary, by his wife Marie Dorothea, duchess of Württemberg. This princess, who was a great-granddaughter of the empress Maria Theresa, and a great-niece of Marie Antoinette, endeared herself to the people by her elevated character and indefatigable benevolence, while her beauty gained for her the sobriquet of “The Rose of Brabant”; she was also an accomplished artist and musician, and a fine horsewoman. Between the years 1854 and 1865 Leopold travelled much abroad, visiting India and China as well as Egypt and the countries on the Mediterranean coast of Africa. On the 10th of December 1865 he succeeded his father. On the 28th of January 1869 he lost his only son, Leopold (b. 1859), duke of Hainaut. The king’s brother Philip, count of Flanders (1837-1905), then became heir to the throne; and on his death his son Albert (b. 1875) became heir-presumptive. During the Franco-Prussian War (1870-1871) the king of the Belgians preserved neutrality in a period of unusual difficulty and danger. But the most notable event in Leopold’s career was the foundation of the Congo Free State (q.v.). While still duke of Brabant he had been the first to call the attention of the Belgians to the need of enlarging their horizon beyond sea, and after his accession to the throne he gave the first impulse towards the development of this idea by founding in 1876 theAssociation Internationale Africaine. He enlisted the services of H. M. Stanley, who visited Brussels in 1878 after exploring the Congo river, and returned in 1879 to the Congo as agent of theComité d’Études du Haut Congo, soon afterwards reorganized as the “International Association of the Congo.” This association was, in 1884-1885, recognized by the powers as a sovereign state under the name of theÉtat Indépendant du Congo. Leopold’s exploitation of this vast territory, which he administered autocratically, and in which he associated himself personally with various financial schemes, was understood to bring him an enormous fortune; it was the subject of acutely hostile criticism, to a large extent substantiated by the report of a commission of inquiry instituted by the king himself in 1904, and followed in 1908 by the annexation of the state to Belgium (seeCongo Free State:History). In 1880 Leopold sought an interview with General C. G. Gordon and obtained his promise, subject to the approval of the British government, to enter the Belgian service on the Congo. Three years later Leopold claimed fulfilment of the promise, and Gordon was about to proceed to the Congo when the British government required his services for the Sudan. On the 15th of November 1902 King Leopold’s life was attempted in Brussels by an Italian anarchist named Rubino. Queen Marie Henriette died at Spa on the 19th of September of the same year. Besides the son already mentioned she had borne to Leopold three daughters—Louise Marie Amélie (b. 1858), who in 1875 married Philip of Saxe-Coburg and Gotha, and was divorced in 1906; Stéphanie (b. 1864), who married Rudolph, crown prince of Austria, in 1881, and after his death in 1889 married, against her father’s wishes, Elemer, Count Lonyay, in 1900; and Clémentine (b. 1872). At the time of the queen’s death an unseemly incident was occasioned by Leopold’s refusal to see his daughter Stéphanie, who in consequence was not present at her mother’s funeral. The disagreeable impression on the public mind thus created was deepened by an unfortunate litigation, lasting for two years (1904-1906), over the deceased queen’s will, in which the creditors of the princess Louise, together with princess Stéphanie (Countess Lonyay), claimed that under the Belgian law the queen’s estate was entitled to half of her husband’s property. This claim was disallowed by the Belgian courts. The king died at Laeken, near Brussels, on the 17th of December 1909. On the 23rd of that month his nephew took the oath to observe the constitution, assuming the title of Albert I. King Leopold was personally a man of considerable attainments and much strength of character, but he was a notoriously dissolute monarch, who even to the last offended decent opinion by his indulgences at Paris and on the Riviera. The wealth he amassed from the Congo he spent, no doubt, royally not only in this way but also on public improvements in Belgium; but he had a hard heart towards the natives of his distant possession.

LEOPOLD II.(1797-1870), of Habsburg-Lorraine, grand-duke of Tuscany, was born on the 3rd of October 1797, the son of the grand-duke Ferdinand III., whom he succeeded in 1824. During the first twenty years of his reign he devoted himself to the internal development of the state. His was the mildest and least reactionary of all the Italian despotisms of the day, and although always subject to Austrian influence he refused to adopt the Austrian methods of government, allowed a fair measure of liberty to the press, and permitted many political exiles from other states to dwell in Tuscany undisturbed. But when in the early ’forties a feeling of unrest spread throughout Italy, even in Tuscany demands for a constitution and other political reforms were advanced; in 1845-1846 riots broke out in various parts of the country, and Leopold granted a number of administrative reforms. But Austrian influence prevented him from going further, even had he wished to do so. The election of Pope Pius IX. gave fresh impulse to the Liberal movement, and on the 4th of September 1847 Leopold instituted the National Guard—a first step towards the constitution; shortly after the marchese Cosimo Ridolfi was appointed prime minister. The granting of the Neapolitan and Piedmontese constitutions was followed (17th February 1848) by that of Tuscany, drawn up by Gino Capponi. The revolution in Milan and Vienna aroused a fever of patriotic enthusiasm in Tuscany, where war against Austria was demanded; Leopold, giving way to popular pressure, sent a force of regulars and volunteers to co-operate with Piedmont in the Lombard campaign. His speech on their departure was uncompromisingly Italian and Liberal. “Soldiers,” he said, “the holy cause of Italian freedom is being decided to-day on the fields of Lombardy. Already the citizens of Milan have purchased their liberty with their blood and with a heroism of which history offers few examples.... Honour to the arms of Italy! Long live Italian independence!” The Tuscan contingent fought bravely, if unsuccessfully, at Curtatone and Montanara. On the 26th of June the first Tuscan parliament assembled, but thedisturbances consequent on the failure of the campaign in Lombardy led to the resignation of the Ridolfi ministry, which was succeeded by that of Gino Capponi. The riots continued, especially at Leghorn, which was a prey to actual civil war, and the democratic party of which F. D. Guerrazzi and G. Montanelli were leading lights became every day more influential. Capponi resigned, and Leopold reluctantly agreed to a Montanelli-Guerrazzi ministry, which in its turn had to fight against the extreme republican party. New elections in the autumn of 1848 returned a constitutional majority, but it ended by voting in favour of a constituent assembly. There was talk of instituting a central Italian kingdom with Leopold as king, to form part of a larger Italian federation, but in the meanwhile the grand-duke, alarmed at the revolutionary and republican agitations in Tuscany and encouraged by the success of the Austrian arms, was, according to Montanelli, negotiating with Field-Marshal Radetzky and with Pius IX., who had now abandoned his Liberal tendencies, and fled to Gaeta. Leopold had left Florence for Siena, and eventually for Porto S. Stefano, leaving a letter to Guerrazzi in which, on account of a protest from the pope, he declared that he could not agree to the proposed constituent assembly. The utmost confusion prevailed in Florence and other parts of Tuscany. On the 9th of February 1849 the republic was proclaimed, largely as a result of Mazzini’s exhortations, and on the 18th Leopold sailed for Gaeta. A third parliament was elected and Guerrazzi appointed dictator. But there was great discontent, and the defeat of Charles Albert at Novara caused consternation among the Liberals. The majority, while fearing an Austrian invasion, desired the return of the grand-duke who had never been unpopular, and in April 1849 the municipal council usurped the powers of the assembly and invited him to return, “to save us by means of the restoration of the constitutional monarchy surrounded by popular institutions, from the shame and ruin of a foreign invasion.” Leopold accepted, although he said nothing about the foreign invasion, and on the 1st of May sent Count Luigi Serristori to Tuscany with full powers. But at the same time the Austrians occupied Lucca and Leghorn, and although Leopold simulated surprise at their action it has since been proved, as the Austrian general d’Aspre declared at the time, that Austrian intervention was due to the request of the grand-duke. On the 24th of May the latter appointed G. Baldasseroni prime minister, on the 25th the Austrians entered Florence and on the 28th of July Leopold himself returned. In April 1850 he concluded a treaty with Austria sanctioning the continuation for an indefinite period of the Austrian occupation with 10,000 men; in September he dismissed parliament, and the following year established a concordat with the Church of a very clerical character. He feebly asked Austria if he might maintain the constitution, and the Austrian premier, Prince Schwarzenberg, advised him to consult the pope, the king of Naples and the dukes of Parma and Modena. On their advice he formally revoked the constitution (1852). Political trials were held, Guerrazzi and many others being condemned to long terms of imprisonment, and although in 1855 the Austrian troops left Tuscany, Leopold’s popularity was gone. A part of the Liberals, however, still believed in the possibility of a constitutional grand-duke who could be induced for a second time to join Piedmont in a war against Austria, whereas the popular party headed by F. Bartolommei and G. Dolfi realized that only by the expulsion of Leopold could the national aspirations be realized. When in 1859 France and Piedmont made war on Austria, Leopold’s government failed to prevent numbers of young Tuscan volunteers from joining the Franco-Piedmontese forces. Finally an agreement was arrived at between the aristocratic constitutionalists and the popular party, as a result of which the grand-duke’s participation in the war was formally demanded. Leopold at first gave way, and entrusted Don Neri Corsini with the formation of a ministry. The popular demands presented by Corsini were for the abdication of Leopold in favour of his son, an alliance with Piedmont and the reorganization of Tuscany in accordance with the eventual and definite reorganization of Italy. Leopold hesitated and finally rejected the proposals as derogatory to his dignity. On the 27th of April there was great excitement in Florence, Italian colours appeared everywhere, but order was maintained, and the grand-duke and his family departed for Bologna undisturbed. Thus the revolution was accomplished without a drop of blood being shed, and after a period of provisional government Tuscany was incorporated in the kingdom of Italy. On the 21st of July Leopold abdicated in favour of his son Ferdinand IV., who never reigned, but issued a protest from Dresden (26th March 1860). He spent his last years in Austria, and died in Rome on the 29th of January 1870.

Leopold of Tuscany was a well-meaning, not unkindly man, and fonder of his subjects than were the other Italian despots, but he was weak, and too closely bound by family ties and Habsburg traditions ever to become a real Liberal. Had he not joined the conclave of autocrats at Gaeta, and, above all, had he not summoned Austrian assistance while denying that he had done so, in 1849, he might yet have preserved his throne, and even changed the whole course of Italian history. At the same time his rule, if not harsh, was enervating and demoralizing.

See G. Baldasseroni,Leopoldo II.(Florence, 1871), useful but reactionary in tendency, the author having been Leopold’s minister, G. Montanelli,Memorie sull’ Italia(Turin, 1853); F. D. Guerrazzi,Memorie(Leghorn, 1848); Zobi,Storia civile della Toscana, vols. iv.-v. (Florence, 1850-1852); A. von Reumont,Geschichte Toscanas(2 vols., Gotha, 1876-1877); M. Bartolommei-Gioli,Il Rivolgimento Toscano e L’azione popolare(Florence, 1905); C. Tivaroni,L’ Italia durante il dominio Austriaco, vol. i. (Turin, 1892), andL’ Italia degli Italiani, vol. i. (Turin, 1895). See alsoRicasoli;Bartolommei;Capponi, Gino; &c.

See G. Baldasseroni,Leopoldo II.(Florence, 1871), useful but reactionary in tendency, the author having been Leopold’s minister, G. Montanelli,Memorie sull’ Italia(Turin, 1853); F. D. Guerrazzi,Memorie(Leghorn, 1848); Zobi,Storia civile della Toscana, vols. iv.-v. (Florence, 1850-1852); A. von Reumont,Geschichte Toscanas(2 vols., Gotha, 1876-1877); M. Bartolommei-Gioli,Il Rivolgimento Toscano e L’azione popolare(Florence, 1905); C. Tivaroni,L’ Italia durante il dominio Austriaco, vol. i. (Turin, 1892), andL’ Italia degli Italiani, vol. i. (Turin, 1895). See alsoRicasoli;Bartolommei;Capponi, Gino; &c.

(L. V.*)

LEOPOLD II., a lake of Central Africa in the basin of the Kasai affluent of the Congo, cut by 2° S. and 18° 10′ E. It has a length N. to S. of about 75 m., is 30 m. across at its northern end, tapering towards its southern end. Numerous bays and gulfs render its outline highly irregular. Its shores are flat and marshy, the lake being (in all probability) simply the lowest part of a vast lake which existed here before the Kasai system breached the barrier—at Kwa mouth—separating it from the Congo. The lake is fed by the Lokoro (about 300 m. long) and smaller streams from the east. Its northern and western affluents are comparatively unimportant. It discharges its waters (at its southern end) into the Mfini, which is in reality the lower course of the Lukenye. The lake is gradually diminishing in area; in the rainy season it overflows its banks. The surrounding country is very flat and densely wooded.

SeeKasai; and articles and maps inLe Mouvement géog., specially vol. xiv., No. 29 (1897) and vol. xxiv., No. 38 (1907).

SeeKasai; and articles and maps inLe Mouvement géog., specially vol. xiv., No. 29 (1897) and vol. xxiv., No. 38 (1907).

LEOTYCHIDES,Spartan king, of the Eurypontid family, was descended from Theopompus through his younger son Anaxandridas (Herod. viii. 131), and in 491B.C.succeeded Demaratus (q.v.), whose title to the throne he had with Cleomenes’ aid successfully challenged. He took part in Cleomenes’ second expedition to Aegina, on which ten hostages were seized and handed over to the Athenians for safe custody: for this he narrowly escaped being surrendered to the Aeginetans after Cleomenes’ death. In the spring of 479 we find him in command of the Greek fleet of 110 ships, first at Aegina and afterwards at Delos. In August he attacked the Persian position at Mycale on the coast of Asia Minor opposite Samos, inflicted a crushing defeat on the land-army, and annihilated the fleet which was drawn up on the shore. Soon afterwards he sailed home with the Peloponnesians, leaving the Athenians to prosecute the siege of Sestos. In 476 he led an army to Thessaly to punish the Aleuadae of Larisa for the aid they had rendered to the Persians and to strengthen Spartan influence in northern Greece. After a series of successful engagements he accepted a bribe from the enemy to withdraw. For this he was brought to trial at Sparta, and to save his life fled to the temple of Athena Alea at Tegea. Sentence of exile was passed, his house was razed and his grandson Archidamus II. ascended the throne (Herod. vi. 65-87, ix. 90-114; Thucydides i. 89; Pausanias iii. 4. 3. 7. 9-10; Plutarch,De malignitate Herodoti, 21, p. 859D; Diodorus xi. 34-37).

According to Diodorus (xi. 48) Leotychides reigned twenty-two, his successor Archidamus forty-two years. The total duration of the two reigns, sixty-four years, we know to be correct, for Leotychides came to the throne in 491 and Archidamus (q.v.) died in 427. On this basis, then, Leotychides’s exile would fall in 469 and the Thessalian expedition in that or the preceding year (so E. Meyer,Geschichte des Altertums, iii. § 287). But Diodorus is not consistent with himself; he attributes (xi. 48) Leotychides’s death to the year 476-475 and he records (xii. 35) Archidamus’s death in 434-433, though he introduces him in the following years at the head of the Peloponnesian army (xii. 42, 47, 52). Further, he says expressly that Leotychidesἐτελεύτησεν ἄρξας ἔτη εἴκοσι καὶ δύο,i.e.he lived twenty-two years after his accession. The twenty-two years, then, may include the time which elapsed between his exile and his death. In that case Leotychides died in 469, and 476-475 may be the year in which his reign, though not his life, ended. This date seems, from what we know of the political situation in general, to be more probable than the later one for the Thessalian campaign.G. Busolt,Griech. Geschichte, iii. 83, note; J. B. Bury,History of Greece, p. 326; G. Grote,History of Greece, new edition 1888, iv. 349, note; also abridged edition 1907, p. 273, note 3. Beloch’s view (Griech. Geschichte, i. 455, note 2) that the expedition took place in 476, the trial and flight in 469, is not generally accepted.

According to Diodorus (xi. 48) Leotychides reigned twenty-two, his successor Archidamus forty-two years. The total duration of the two reigns, sixty-four years, we know to be correct, for Leotychides came to the throne in 491 and Archidamus (q.v.) died in 427. On this basis, then, Leotychides’s exile would fall in 469 and the Thessalian expedition in that or the preceding year (so E. Meyer,Geschichte des Altertums, iii. § 287). But Diodorus is not consistent with himself; he attributes (xi. 48) Leotychides’s death to the year 476-475 and he records (xii. 35) Archidamus’s death in 434-433, though he introduces him in the following years at the head of the Peloponnesian army (xii. 42, 47, 52). Further, he says expressly that Leotychidesἐτελεύτησεν ἄρξας ἔτη εἴκοσι καὶ δύο,i.e.he lived twenty-two years after his accession. The twenty-two years, then, may include the time which elapsed between his exile and his death. In that case Leotychides died in 469, and 476-475 may be the year in which his reign, though not his life, ended. This date seems, from what we know of the political situation in general, to be more probable than the later one for the Thessalian campaign.

G. Busolt,Griech. Geschichte, iii. 83, note; J. B. Bury,History of Greece, p. 326; G. Grote,History of Greece, new edition 1888, iv. 349, note; also abridged edition 1907, p. 273, note 3. Beloch’s view (Griech. Geschichte, i. 455, note 2) that the expedition took place in 476, the trial and flight in 469, is not generally accepted.

(M. N. T.)

LEOVIGILD,orLöwenheld(d. 586), king of the Visigoths, became king in 568 after the short period of anarchy which followed the death of King Athanagild, whose widow, Goisvintha, he married. At first he ruled that part of the Visigothic kingdom which lay to the south of the Pyrenees, his brother Liuva or Leova governing the small part to the north of these mountains; but in 572 Liuva died and Leovigild became sole king. At this time the Visigoths who settled in Spain early in the 5th century were menaced by two powerful enemies, the Suevi who had a small kingdom in the north-west of the peninsula, and the Byzantines who had answered Athanagild’s appeal for help by taking possession of a stretch of country in the south-east. Their kingdom, too, was divided and weakened by the fierce hostility between the orthodox Christians and those who professed Arianism. Internal and external dangers alike, however, failed to daunt Leovigild, who may fairly be called the restorer of the Visigothic kingdom. He turned first against the Byzantines, who were defeated several times; he took Cordova and chastised the Suevi; and then by stern measures he destroyed the power of those unruly and rebellious chieftains who had reduced former kings to the position of ciphers. The chronicler tells how, having given peace to his people, he, first of the Visigothic sovereigns, assumed the attire of a king and made Toledo his capital. He strengthened the position of his family and provided for the security of his kingdom by associating his two sons, Recared and Hermenegild, with himself in the kingly office and placing parts of the land under their rule. Leovigild himself was an Arian, being the last of the Visigothic kings to hold that creed; but he was not a bitter foe of the orthodox Christians, although he was obliged to punish them when they conspired against him with his external enemies. His son Hermenegild, however, was converted to the orthodox faith through the influence of his Frankish wife, Ingundis, daughter of King Sigebert I., and of Leander, metropolitan of Seville. Allying himself with the Byzantines and other enemies of the Visigoths, and supported by most of the orthodox Christians he headed a formidable insurrection. The struggle was fierce; but at length, employing persuasion as well as force, the old king triumphed. Hermenegild was captured; he refused to give up his faith and in March or April 585 he was executed. He was canonized at the request of Philip II., king of Spain, by Pope Sixtus V. About this time Leovigild put an end to the kingdom of the Suevi. During his last years he was engaged in a war with the Franks. He died at Toledo on the 21st of April 586 and was succeeded by his son Recared.

LEPANTO,1BATTLE OF,fought on the 7th of October 1571. The conquest of Cyprus by the Turks, and their aggressions on the Christian powers, frightened the states of the Mediterranean into forming a holy league for their common defence. The main promoter of the league was Pope Pius V., but the bulk of the forces was supplied by the republic of Venice and Philip II. of Spain, who was peculiarly interested in checking the Turks both because of the Moorish element in the population of Spain, and because he was also sovereign of Naples and Sicily. In compliment to King Philip, the general command of the league’s fleet was given to his natural brother, Don John of Austria. It included, however, only twenty-four Spanish ships. The great majority of the two hundred galleys and eight galeasses, of which the fleet was composed, came from Venice, under the command of the proveditore Barbarigo; from Genoa, which was in close alliance with Spain, under Gianandrea Doria; and from the Pope whose squadron was commanded by Marc Antonio Colonna. The Sicilian and Neapolitan contingents were commanded by the marquess of Santa Cruz, and Cardona, Spanish officers. Eight thousand Spanish soldiers were embarked. The allied fleet was collected slowly at Messina, from whence it advanced by the passage between Ithaca and Cephalonia to Cape Marathia near Dragonera. The Turkish fleet which had come up from Cyprus and Crete anchored in the Gulf of Patras. It consisted in all of 273 galleys which were of lighter build than the Christians’, and less well supplied with cannon or small arms. The Turks still relied mainly on the bow and arrow. Ali, the capitan pasha, was commander-in-chief, and he had with him Chulouk Bey of Alexandria, commonly called Scirocco, and Uluch Ali, dey of Algiers. On the 7th of October the Christian fleet advanced to the neighbourhood of Cape Scropha. It was formed in the traditional order of the galleys—a long line abreast, subdivided into the centre or “battle” commanded by Don John in person, the left wing under the proveditore Barbarigo, and the right under Gianandrea Doria. But a reserve squadron was placed behind the centre under the marquess of Santa Cruz, and the eight lumbering galeasses were stationed at intervals in front of the line to break the formation of the Turks. The capitan pasha left his anchorage in the Gulf of Patras with his fleet in a single line, without reserve or advance-guard. He was himself in the centre, with Scirocco on his right and Uluch Ali on his left. The two fleets met south of Cape Scropha, both drawn up from north to south, the land being close to the left flank of the Christians, and the right of the Turks. To the left of the Turks and the right of the Christians, there was open sea. Ali Pasha’s greater numbers enabled him to outflank his enemy. The Turks charged through the intervals between the galeasses, which proved to be of no value. On their right Scirocco outflanked the Venetians of Barbarigo, but the better build of the galleys of Saint Mark and the admirable discipline of their crews gave them the victory. The Turks were almost all sunk or driven on shore. Scirocco and Barbarigo both lost their lives. On the centre Don John and the capitan pasha met prow to prow—the Christians reserving the fire of their bow guns (calleddi cursia) till the moment of impact, and then boarding. Ali Pasha was slain and his galley taken. Everywhere on the centre the Christians gained the upper hand, but their victory was almost turned into a defeat by the mistaken manœuvres of Doria. In fear lest he should be outflanked by Uluch Ali, he stood out to sea, leaving a gap between himself and the centre. The dey of Algiers, who saw the opening, reversed the order of his squadron, and fell on the right of the centre. The galleys of the Order of Malta, which were stationed at this point, suffered severely, and their flagship was taken with great slaughter. A disaster was averted by the marquess of Santa Cruz, who brought up the reserve. Uluch Ali then retreated with sail and oar, bringing most of his division off in good order.

The loss of life in the battle was enormous, being put at 20,000 for the Turks and 8000 for the Christians. The battle of Lepanto was of immense political importance. It gave the naval power of the Turks a blow from which it never recovered, and put a stop to their aggression in the Eastern Mediterranean. Historically the battle is interesting because it was the last example of an encounter on a great scale between fleets of galleys and also because it was the last crusade. The Christian powers of the Mediterranean did really combine to avert the ruin of Christendom. Hardly a noble house of Spain or Italy was not represented in the fleet, and the princes headed the boarders. Volunteers came from all parts of Europe, and it is said thatamong them was Sir Richard Grenville, afterwards famous for his fight in the “Revenge” off Flores in the Azores. Cervantes was undoubtedly present, and had his left hand shattered by a Turkish bullet.

For full accounts of the battle, with copious references to authorities and to ancient controversies, mostly arising out of the conduct of Doria, see Sir W. Stirling Maxwell,Don John of Austria(1883); and Jurien de la Gravière,La Guerre de Chypre et la bataille de Lepanto(1888).

For full accounts of the battle, with copious references to authorities and to ancient controversies, mostly arising out of the conduct of Doria, see Sir W. Stirling Maxwell,Don John of Austria(1883); and Jurien de la Gravière,La Guerre de Chypre et la bataille de Lepanto(1888).

(D. H.)

1For Lepanto seeNaupactus.

1For Lepanto seeNaupactus.

LE PAUTRE, JEAN(1618-1682), French designer and engraver. He was apprenticed to a carpenter and builder and in addition to learning mechanical and constructive work developed considerable facility with the pencil. His designs, which were innumerable in quantity and exuberant in fancy, consisted mainly of ceilings, friezes, chimney-pieces, doorways and mural decorations; he also devised fire-dogs, sideboards, cabinets, console tables, mirrors and other pieces of furniture; he was long employed at the Gobelins. His work is often excessively flamboyant and over-elaborate; he revelled in amorini and swags, arabesques and cartouches. His chimney-pieces, however, were frequently simple and elegant. His engraved plates, almost entirely original, are something like 1500 in number and include a portrait of himself. He became a member of the academy of Paris in 1677.

LEPCHA,the name of the aboriginal inhabitants of Sikkim (q.v.). A peace-loving people, the Lepchas have been repeatedly conquered by surrounding hill-tribes, and their ancient patriarchal customs are dying out. The total number of speakers of Lepcha, or Rong, in all India in 1901, was only 19,291. Their rich and beautiful language has been preserved from extinction by the efforts of General Mainwaring and others; but their literature was almost entirely destroyed by the Tibetans, and their traditions are being rapidly forgotten. Once free and independent, they are now the poorest people in Sikkim, and it is from them that the coolie class is drawn. They are above all things woodmen, knowing the ways of beasts and birds, and possessing an extensive zoological and botanical nomenclature of their own.

See Florence Donaldson,Lepcha Land(1900).

See Florence Donaldson,Lepcha Land(1900).

LE PELETIER(orLepelletier),DE SAINT-FARGEAU, LOUIS MICHEL(1760-1793), French politician, was born on the 29th of May 1760 at Paris. He belonged to a well-known family, his great-grandfather, Michel Robert Le Peletier des Forts, count of Saint-Fargeau, having been controller-general of finance. He inherited a great fortune, and soon became president of the parlement of Paris and in 1789 he was a deputy of thenoblesseto the States-General. At this time he shared the conservative views of the majority of his class; but by slow degrees his ideas changed and became very advanced. On the 13th of July 1789 he demanded the recall of Necker, whose dismissal by the king had aroused great excitement in Paris; and in the Constituent Assembly he had moved the abolition of the penalty of death, of the galleys and of branding, and the substitution of beheading for hanging. This attitude won him great popularity, and on the 21st of June 1790 he was made president of the Constituent Assembly. During the existence of the Legislative Assembly, he was president of the general council for the department of the Yonne, and was afterwards elected by this department as a deputy to the Convention. Here he was in favour of the trial of Louis XVI. by the assembly and voted for the death of the king. This vote, together with his ideas in general, won him the hatred of the royalists, and on the 20th of January 1793, the eve of the execution of the king, he was assassinated in the Palais Royal at Paris by a member of the king’s body-guard. The Convention honoured Le Peletier by a magnificent funeral, and the painter J. L. David represented his death in a famous picture, which was later destroyed by his daughter. Towards the end of his life, Le Peletier had interested himself in the question of public education; he left fragments of a plan, the ideas contained in which were borrowed in later schemes. His assassin fled to Normandy, where, on the point of being discovered, he blew out his brains. Le Peletier had a brother, Félix (1769-1837), well known for his advanced ideas. His daughter, Suzanne Louise, was “adopted” by the French nation.

SeeŒuvres de M. le Peletier de Saint-Fargeau(Brussels, 1826) with a life by his brother Félix; E. Le Blant, “Le Peletier de St-Fargeau, et son meurtrier,” in theCorrespondantreview (1874); F. Clerembray,Épisodes de la Révolution(Rouen, 1891); Brette, “La Réforme de la législation universelle, et le plan de Lepelletier Saint-Fargeau,” inLa Révolution française, xlii. (1902); and M. Tourneux,Bibliog. de l’hist. de Paris ...(vol. i., 1890, Nos. 3896-3910, and vol. iv., 1906,s.v.Lepeletier).

SeeŒuvres de M. le Peletier de Saint-Fargeau(Brussels, 1826) with a life by his brother Félix; E. Le Blant, “Le Peletier de St-Fargeau, et son meurtrier,” in theCorrespondantreview (1874); F. Clerembray,Épisodes de la Révolution(Rouen, 1891); Brette, “La Réforme de la législation universelle, et le plan de Lepelletier Saint-Fargeau,” inLa Révolution française, xlii. (1902); and M. Tourneux,Bibliog. de l’hist. de Paris ...(vol. i., 1890, Nos. 3896-3910, and vol. iv., 1906,s.v.Lepeletier).

LEPIDOLITE, orLithia-Mica, a mineral of the mica group (seeMica). It is a basic aluminium, potassium and lithium fluo-silicate, with the approximate formula KLi [Al(OH, F)2] Al(SiO3)3. Lithia and fluorine are each present to the extent of about 5%; rubidium and caesium are sometimes present in small amounts. Distinctly developed monoclinic crystals or cleavage sheets of large size are of rare occurrence, the mineral being usually found as scaly aggregates, and on this account was named lepidolite (from Gr.λεπίς, scale) by M. H. Klaproth in 1792. It is usually of a lilac or peach-blossom colour, but is sometimes greyish-white, and has a pearly lustre on the cleavage surfaces. The hardness is 2½-4 and the sp. gr. 2.8-2.9, the optic axial angle measures 50°-70°. It is found in pegmatite-veins, often in association with pink tourmaline (rubellite) and sometimes intergrown in parallel position with muscovite. Scaly masses of considerable extent are found at Rozena near Bystrzitz in Moravia and at Pala in San Diego county, California. The material from Rozena has been known since 1791, and has sometimes been cut and polished for ornamental purposes: it has a pretty colour and spangled appearance and takes a good polish, but is rather soft. At Pala it has been extensively mined for the preparation of lithium and rubidium salts. Other localities for the mineral are the island of Utö in Sweden, and Auburn and Paris in Maine, U.S.A.; at Alabashka near Mursinka in the Urals large isolated crystals have been found, and from Central Australia transparent cleavage sheets of a fine lilac colour are known.

The lithium-iron micazinnwalditeorlithioniteis closely allied to lepidolite, differing from it in containing some ferrous iron in addition to the constituents mentioned above. It occurs as greyish silvery scales with hexagonal outlines in the tin-bearing granites of Zinnwald in the Erzgebirge, Bohemia and of Cornwall.

(L. J. S.)

LEPIDOPTERA(Gr.λεπίς, a scale or husk, andπτερόν, a wing), a term used in zoological classification for one of the largest and best-known orders of the class Hexapoda (q.v.), in order that comprises the insects popularly called butterflies and moths. The term was first used by Linnaeus (1735) in the sense still accepted by modern zoologists, and there are few groups of animals as to whose limits and distinguishing characters less controversy has arisen.

Characters.—The name of the order indicates the fact that the wings (and other parts of the body) are clothed with flattenedcuticular structures—the scales (fig. 7)—that may be regarded as modified arthropodan “hairs.” Such scales are not peculiar to the Lepidoptera—they are found also on many of the Aptera, on the Psocidae, a family of Corrodentia, on some Coleoptera (beetles) and on the gnats (Culicidae), a family of Diptera. The most distinctive structural features of the Lepidoptera are to be found in the jaws. The mandibles are mere vestiges or entirely absent; the second maxillae are usually reduced to a narrow transverse mentum which bears the scale-covered labial palps, between which project the elongate first maxillae, grooved on their inner faces, so as to form when apposed a tubular proboscis adapted for sucking liquid food.

All Lepidoptera are hatched as the eruciform soft-bodied type of larva (fig. 1,a) known as the caterpillar, with biting mandibles, three pairs of thoracic legs and with a variable number (usually five pairs) of abdominal prolegs, which carry complete or incomplete circles of hooklets. The pupa in a single family only is free (i.e.with the appendages free from the body), and mandibulate. In the vast majority of the order it is more or less obtect (i.e.with the appendages fixed to the cuticle of the body) and without mandibles (fig. 1,c).

a, Palp.

b, Galea.

c, Lacinia.

d, Stipes.

e, Cardo of maxilla.

Structure.—The head in the Lepidoptera is sub-globular in shape with the compound eyes exceedingly well developed, and with a pair of ocelli or “simple eyes” often present on the vertex. It is connected to the thorax by a relatively broad and membranous “neck.” The feelers are many-jointed, often they are complex, the segments bearing processes arranged in a comb-like manner and furnished with numerous sensory hairs (fig. 2). The complexity of the feelers is carried to its highest development in certain male moths that have a wonderful power of discovering their females by smell or some analogous sense. Often the feelers are excessively complex in male moths whose maxillae are so reduced that they take no food in the imaginal state. The nature of the jaws has already been briefly described. Functional mandibles of peculiar form (fig. 3, A) are present in the remarkable small moths of the genusMicropteryx(orEriocephala), and there are vestiges of these jaws in other moths of low type, but the minute structures in the higher Lepidoptera that were formerly described as mandibles are now believed to belong to the labrum, the true mandibles being perhaps represented by rounded prominences, not articulated with the head-capsule. Throughout the order, as a whole, the jaws are adapted for sucking liquid food, and the suctorial proboscis (often erroneously called a “tongue”) is formed as was shown by J. C. Savigny in 1816 by two elongated and flexible outgrowths of the first maxillae, usually regarded as representing the outer lobes or galeae (fig. 4, A, B,g). These structures are grooved along their inner faces and by means of a series of interlocking hair-like bristles can be joined together so as to form a tubular sucker (fig. 4, C). At their extremities they are beset with club-like sense-organs, whose apparent function is that of taste. The proboscis when in use is stretched out in front of the head and inserted into the corolla of a flower or elsewhere, for the absorption of liquid nourishment. When at rest, the proboscis is rolled up into a close spiral beneath the head and between the labial palps (fig. 4, A,p). Only in the genusMicropteryxmentioned above is the lacinia of the maxilla (as A. Walter has shown) developed (fig. 3, B,c). The maxillary palp is usually a mere vestige (fig. 4, B,p) though it is conspicuous in a few families of small moths. A considerable number of Lepidoptera take no food in the imaginal state; in these the maxillae are reduced or altogether atrophied. The second maxillae are intimately fused together to form the labium, which consists only of a reduced mentum, bearing sometimes vestigial lobes and always a pair of palps. These have two or three segments and are clothed with scales. The form and direction of the terminal segment of the labial palp afford valuable characters in classification.Fig. 4.—Arrangement of the jaws in a typical Moth. Somewhat diagrammatic and in part after E. Burgess and V. L. Kellogg (Amer. Nat.xiv. xxix.).A, Front view of head.c, Clypeus.e, Compound eye.m, Vestigial mandible.l, Labrum.g, Galeae of 1st maxillae.p, Labial palp. Magnified, B.b, Base of first maxilla dissected out of the head.p, Vestigial palp.g, Galea. Further magnified.C, Part transverse section showing how the channel (A) of the proboscis is formed by the interlocking of the grooved inner faces of the flexible maxillae.t, Air-tube.n, Nerve.m, Muscle-fibres. Highly magnified.In the thorax of the Lepidoptera the foremost segment or prothorax is very small, and not movable on the mesothorax. In many families it carries a pair of small erectile plates—the patagia—which have been regarded as serially homologous with the wings. The mesothorax is extensive; its scutum forming most of the dorsal thoracic area and small plates—tegulae—are often present at the base of the forewings, as in Hymenoptera. The tegulae which are beset with long hair-like scales are often conspicuous. The metathorax is smaller than the mesothorax. The legs are of the typical hexapodan form with five-segmented feet; the shins often bear terminal and median spurs articulated at their bases and the entire limbs are clothed with scales.After A. S. Packard,Mem. Nat. Acad. Sci.vol. vii.Fig. 5.—Wing-neuration of a Notodont Moth. 2, Sub-costal; 3, radial; 4, median; 5, cubital; 7, 8, anal nervures.a, Discoidal areolet or “cell”;f, frenulum. Note that the forewing has five branches (1-5) of the radial nervure, the hindwing one only. The first anal nervure (No. 6) is absent.The wings of the Lepidoptera may be said to dominate the structure of the insect; only exceptionally, in certain female moths, are they vestigial or absent (fig. 17). The forewing, with its prominent apex, is longer than the hindwing, and the neuration in both (see figs. 5 and 6) is for the most part longitudinal, only a few transverse nervures, which are, in fact, branches of the median trunk, marking off a discoidal areolet or “cell” (fig. 5,a). The five branches of the radial nervure (figs. 5, 6,3) (seeHexapoda) are usually present in the forewing, but the hindwing, in most families, has only a single radial nervure; its anal area is, however, often more strongly developed than that of the forewing. The two wings of a side are usually kept together during flight by a few stout bristles—the frenulum—(fig. 5,f) projecting from the base of the costa of the hindwing and fitting beneath a membranous fold or a few thickened scales—the retinaculum—on the under surface of the forewing. In butterflies there is no frenulum, but a costal outgrowth of thehindwing subserves the same function. In the most primitive moths a small lobate outgrowth—the jugum (fig. 6,j.)—from the dorsum of the forewing is present, but it can be of little service in keeping the two wings together. A jugum may be also present on the hindwing. The legs, which are generally used for clinging rather than for walking, have five-segmented feet and are covered with scales. In some families the front pair are reduced and without tarsal segments.After Packard,Mem. Nat. Acad. Sci.vol. vii.Fig. 6.—Wing neuration of a Swift Moth (Hepialid).j, Jugum. Nervures numbered as in fig. 5. Note that there are five branches to the radial nervure (No. 3) in both fore- and hindwing, and that the median trunk nervures (No. 4) traverse the discoidal areolet.Fig. 7.—A, Arrangement of scales in rows on wing of Butterfly.n, Nervure;c, collar-like outgrowths of cuticle. Magnified. B, single scale, and C, an androconium more highly magnified.Ten abdominal segments are recognizable in many Lepidoptera, but the terminal segments are reduced or modified to form external organs of reproduction. In the male, according to the interpretation of C. Peytoureau, the lateral plates belonging to the ninth segment form paired claspers beset with harpes, or series of ridges or teeth, while the tergum of the tenth segment forms a dorsal hook—the uncus—and its sternum a ventral process or scaphium. In the female the terminal segments form, in some cases, a protrusible ovipositor, but the typical hexapodan ovipositor with its three pairs of processes is undeveloped in the Lepidoptera.As already mentioned, the characteristic scales on the wings, legs and body of the Lepidoptera are cuticular structures. A complete series of transitional forms can be traced between the most elaborate flattened scales (fig. 7, B) with numerous longitudinal striae and a simple arthropod “hair.” Either a “hair” or a scale owes its origin to a special cell of the ectoderm (hypodermis), a process from which grows through the general cuticle and forms around itself the substance of the cuticular appendage. The scales on the wings are arranged in regular rows (fig. 7, A), and the general cuticle is drawn out into a narrow neck or collar around the base of each scale. The scales can be easily rubbed from the surface of the wing, and the series of collars in which the scales rest are then evident (fig. 7, A,c) on the wing-membrane. On the wings of many male butterflies there are specially modified scales—the androconia (fig. 7, C)—which are formed by glandular cells and diffuse a scented secretion. In some cases, the androconia are mixed among the ordinary scales; in others they are associated into conspicuous “brands” (see fig. 66). The admirable colours of the wings of the Lepidoptera are due partly to pigment in the scales—as in the case of yellows, browns, reds and blacks—partly to “interference” effects from the fine striae on the scales—as with the blues, purples and greens.A few points of interest in the internal structure of the Lepidoptera deserve mention. The mouth opens into a sub-globular, muscular pharynx which is believed to suck the liquid food through the proboscis, and force it along the slender gullet into a crop-like enlargement or diverticulum of the fore-gut known as a “food-reservoir” or “sucking-stomach.” The true stomach is tubular, and beyond it lies the intestine into which open the three pairs of excretory (Malpighian) tubes. The terminal part of the intestine is of wide diameter, and in some cases gives off a short caecum. The brain and the sub-oesophageal ganglia are closely approximated; there are two or three thoracic and four (rarely five) abdominal ganglia. In the female each ovary has four ovarian tubes, in which the large egg-cells are enclosed in follicles and associated with nutritive cells. There is a special bursa which in the Hepialidae opens with the vagina on the eighth abdominal sternum. In the Micropterygidae, Enocraniidae and the lower Tineides, the duct of the bursa leads into the vagina, which still opens on the eighth sternum. But in most Lepidoptera, the bursa opens by a vestibule on the eighth sternum, distinct from the vagina, whose opening shifts back to the ninth, the duct of the bursa being connected with the vagina by a canal which opens opposite to the spermatheca. In the male, the two testes are usually fused into a single mass, and a pair of tubular accessory glands open into the vasa deferentia or into the ejaculatory duct. In a few families—the Hepialidae and Saturniidae for example—the testes retain the primitive paired arrangement. These details have been worked out by various students, among whom W. H. Jackson and W. Petersen deserve special mention. Summing up the developmental history of the genital ducts, Jackson remarks that there is “an Ephemeridal stage, which ends towards the close of larval life, an Orthopteran stage, indicated during the quiescent period preceding pupation, and a Lepidopteran stage which begins with the commencement of pupal life.”

Structure.—The head in the Lepidoptera is sub-globular in shape with the compound eyes exceedingly well developed, and with a pair of ocelli or “simple eyes” often present on the vertex. It is connected to the thorax by a relatively broad and membranous “neck.” The feelers are many-jointed, often they are complex, the segments bearing processes arranged in a comb-like manner and furnished with numerous sensory hairs (fig. 2). The complexity of the feelers is carried to its highest development in certain male moths that have a wonderful power of discovering their females by smell or some analogous sense. Often the feelers are excessively complex in male moths whose maxillae are so reduced that they take no food in the imaginal state. The nature of the jaws has already been briefly described. Functional mandibles of peculiar form (fig. 3, A) are present in the remarkable small moths of the genusMicropteryx(orEriocephala), and there are vestiges of these jaws in other moths of low type, but the minute structures in the higher Lepidoptera that were formerly described as mandibles are now believed to belong to the labrum, the true mandibles being perhaps represented by rounded prominences, not articulated with the head-capsule. Throughout the order, as a whole, the jaws are adapted for sucking liquid food, and the suctorial proboscis (often erroneously called a “tongue”) is formed as was shown by J. C. Savigny in 1816 by two elongated and flexible outgrowths of the first maxillae, usually regarded as representing the outer lobes or galeae (fig. 4, A, B,g). These structures are grooved along their inner faces and by means of a series of interlocking hair-like bristles can be joined together so as to form a tubular sucker (fig. 4, C). At their extremities they are beset with club-like sense-organs, whose apparent function is that of taste. The proboscis when in use is stretched out in front of the head and inserted into the corolla of a flower or elsewhere, for the absorption of liquid nourishment. When at rest, the proboscis is rolled up into a close spiral beneath the head and between the labial palps (fig. 4, A,p). Only in the genusMicropteryxmentioned above is the lacinia of the maxilla (as A. Walter has shown) developed (fig. 3, B,c). The maxillary palp is usually a mere vestige (fig. 4, B,p) though it is conspicuous in a few families of small moths. A considerable number of Lepidoptera take no food in the imaginal state; in these the maxillae are reduced or altogether atrophied. The second maxillae are intimately fused together to form the labium, which consists only of a reduced mentum, bearing sometimes vestigial lobes and always a pair of palps. These have two or three segments and are clothed with scales. The form and direction of the terminal segment of the labial palp afford valuable characters in classification.

A, Front view of head.

c, Clypeus.

e, Compound eye.

m, Vestigial mandible.

l, Labrum.

g, Galeae of 1st maxillae.

p, Labial palp. Magnified, B.

b, Base of first maxilla dissected out of the head.

p, Vestigial palp.

g, Galea. Further magnified.

C, Part transverse section showing how the channel (A) of the proboscis is formed by the interlocking of the grooved inner faces of the flexible maxillae.

t, Air-tube.

n, Nerve.

m, Muscle-fibres. Highly magnified.

In the thorax of the Lepidoptera the foremost segment or prothorax is very small, and not movable on the mesothorax. In many families it carries a pair of small erectile plates—the patagia—which have been regarded as serially homologous with the wings. The mesothorax is extensive; its scutum forming most of the dorsal thoracic area and small plates—tegulae—are often present at the base of the forewings, as in Hymenoptera. The tegulae which are beset with long hair-like scales are often conspicuous. The metathorax is smaller than the mesothorax. The legs are of the typical hexapodan form with five-segmented feet; the shins often bear terminal and median spurs articulated at their bases and the entire limbs are clothed with scales.

The wings of the Lepidoptera may be said to dominate the structure of the insect; only exceptionally, in certain female moths, are they vestigial or absent (fig. 17). The forewing, with its prominent apex, is longer than the hindwing, and the neuration in both (see figs. 5 and 6) is for the most part longitudinal, only a few transverse nervures, which are, in fact, branches of the median trunk, marking off a discoidal areolet or “cell” (fig. 5,a). The five branches of the radial nervure (figs. 5, 6,3) (seeHexapoda) are usually present in the forewing, but the hindwing, in most families, has only a single radial nervure; its anal area is, however, often more strongly developed than that of the forewing. The two wings of a side are usually kept together during flight by a few stout bristles—the frenulum—(fig. 5,f) projecting from the base of the costa of the hindwing and fitting beneath a membranous fold or a few thickened scales—the retinaculum—on the under surface of the forewing. In butterflies there is no frenulum, but a costal outgrowth of thehindwing subserves the same function. In the most primitive moths a small lobate outgrowth—the jugum (fig. 6,j.)—from the dorsum of the forewing is present, but it can be of little service in keeping the two wings together. A jugum may be also present on the hindwing. The legs, which are generally used for clinging rather than for walking, have five-segmented feet and are covered with scales. In some families the front pair are reduced and without tarsal segments.

Ten abdominal segments are recognizable in many Lepidoptera, but the terminal segments are reduced or modified to form external organs of reproduction. In the male, according to the interpretation of C. Peytoureau, the lateral plates belonging to the ninth segment form paired claspers beset with harpes, or series of ridges or teeth, while the tergum of the tenth segment forms a dorsal hook—the uncus—and its sternum a ventral process or scaphium. In the female the terminal segments form, in some cases, a protrusible ovipositor, but the typical hexapodan ovipositor with its three pairs of processes is undeveloped in the Lepidoptera.

As already mentioned, the characteristic scales on the wings, legs and body of the Lepidoptera are cuticular structures. A complete series of transitional forms can be traced between the most elaborate flattened scales (fig. 7, B) with numerous longitudinal striae and a simple arthropod “hair.” Either a “hair” or a scale owes its origin to a special cell of the ectoderm (hypodermis), a process from which grows through the general cuticle and forms around itself the substance of the cuticular appendage. The scales on the wings are arranged in regular rows (fig. 7, A), and the general cuticle is drawn out into a narrow neck or collar around the base of each scale. The scales can be easily rubbed from the surface of the wing, and the series of collars in which the scales rest are then evident (fig. 7, A,c) on the wing-membrane. On the wings of many male butterflies there are specially modified scales—the androconia (fig. 7, C)—which are formed by glandular cells and diffuse a scented secretion. In some cases, the androconia are mixed among the ordinary scales; in others they are associated into conspicuous “brands” (see fig. 66). The admirable colours of the wings of the Lepidoptera are due partly to pigment in the scales—as in the case of yellows, browns, reds and blacks—partly to “interference” effects from the fine striae on the scales—as with the blues, purples and greens.

A few points of interest in the internal structure of the Lepidoptera deserve mention. The mouth opens into a sub-globular, muscular pharynx which is believed to suck the liquid food through the proboscis, and force it along the slender gullet into a crop-like enlargement or diverticulum of the fore-gut known as a “food-reservoir” or “sucking-stomach.” The true stomach is tubular, and beyond it lies the intestine into which open the three pairs of excretory (Malpighian) tubes. The terminal part of the intestine is of wide diameter, and in some cases gives off a short caecum. The brain and the sub-oesophageal ganglia are closely approximated; there are two or three thoracic and four (rarely five) abdominal ganglia. In the female each ovary has four ovarian tubes, in which the large egg-cells are enclosed in follicles and associated with nutritive cells. There is a special bursa which in the Hepialidae opens with the vagina on the eighth abdominal sternum. In the Micropterygidae, Enocraniidae and the lower Tineides, the duct of the bursa leads into the vagina, which still opens on the eighth sternum. But in most Lepidoptera, the bursa opens by a vestibule on the eighth sternum, distinct from the vagina, whose opening shifts back to the ninth, the duct of the bursa being connected with the vagina by a canal which opens opposite to the spermatheca. In the male, the two testes are usually fused into a single mass, and a pair of tubular accessory glands open into the vasa deferentia or into the ejaculatory duct. In a few families—the Hepialidae and Saturniidae for example—the testes retain the primitive paired arrangement. These details have been worked out by various students, among whom W. H. Jackson and W. Petersen deserve special mention. Summing up the developmental history of the genital ducts, Jackson remarks that there is “an Ephemeridal stage, which ends towards the close of larval life, an Orthopteran stage, indicated during the quiescent period preceding pupation, and a Lepidopteran stage which begins with the commencement of pupal life.”

Development.—Many observations have been made on the embryology of the Lepidoptera; for some of the more important results of these seeHexapoda. The post-embryonic development of Lepidoptera is more familiar, perhaps, than that of any other group of animals. The egg shows great variation in its outward form, the outer envelope or chorion being in some families globular, in others flattened, in others again erect and sub-conical or cylindrical; while its surface often exhibits a beautifully regular series of ribs and furrows. Throughout the order the larva is of the form known as the caterpillar (fig. 1,a,b, fig. 8B) characterized by the presence of three pairs of jointed and clawed legs on the thorax and a variable number of pairs of abdominal “prolegs”—sub-cylindrical outgrowths of the abdominal segments, provided with a complete or incomplete circle of hooklets at the extremity.

At, Feeler.

Mn, Mandible.

Mx, First maxilla.

Lm, Second maxillae (Labium) with spinneret.


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