2. PLANERA Gmel.

Fig. 282

Leavesobovate-oblong to elliptic, abruptly narrowed at apex into a long point, full and rounded at base on one side and shorter and cuneate on the other, coarsely doubly serrate with slightly incurved teeth, when they unfold coated below with pale pubescence and pilose above with long scattered white hairs, at maturity 4′—6′ long, 1′—3′ wide, dark green and glabrous or scabrate above, pale and soft-pubescent or sometimes glabrous below, with a narrow pale midrib and numerous slender straight primary veins running to the points of the teeth and connected by fine cross veinlets; turning bright clear yellow in the autumn before falling; petioles stout, ¼′ in length; stipules linear-lanceolate, ½′—2′ long.Flowerson long slender drooping pedicels sometimes 1′ in length, in 3 or 4-flowered short-stalkedfascicles; calyx irregularly divided into 7—9 rounded lobes ciliate on the margins, often somewhat oblique, puberulous on the outer surface, green tinged with red above the middle; anthers bright red; ovary light green, ciliate on the margins with long white hairs; styles light green.Fruiton long pedicels in crowded clusters, ripening as the leaves unfold, ovoid to obovoid-oblong, slightly stipitate, conspicuously reticulate-venulose, ½′ long, ciliate on the margins, the sharp points of the wings incurved and inclosing the deep notch.

A tree, sometimes 100°—120° high, with a tall trunk 6°—11° in diameter, frequently enlarged at the base by great buttresses, occasionally rising with a straight undivided shaft to the height of 60°—80° and separating into short spreading branches, more commonly divided 30°—40° from the ground into numerous upright limbs gradually spreading and forming an inversely conic round-topped head of long graceful branches, often 100° or rarely 150° in diameter, and slender branchlets frequently fringing the trunk and its principal divisions, light green and coated at first with soft pale pubescence, becoming in their first winter light reddish brown, glabrous or sometimes puberulous and marked by scattered pale lenticels, and by large elevated semiorbicular leaf-scars showing the ends of three large equidistant fibro-vascular bundles, later becoming dark reddish brown and finally ashy gray.Winter-budsovoid, acute, slightly flattened, about ⅛′ long, with broadly ovate rounded light chestnut-brown glabrous scales, the inner bright green, ovate, acute, becoming on vigorous shoots often nearly 1′ in length.Bark1′—1½′ thick, ashy gray, divided by deep fissures into broad ridges separating on the surface into thin appressed scales.Woodheavy, hard, strong, tough, difficult to split, coarse-grained, light brown, with thick somewhat lighter colored sapwood; largely used for the hubs of wheels, saddle-trees, in flooring and cooperage, and in boat and shipbuilding.

Distribution.River-bottom lands, intervales, low rich hills, and the banks of streams; southern Newfoundland to the northern shores of Lake Superior and the headwaters of the Saskatchewan, southward to the neighborhood of Lake Istokpoga, De Soto County, Florida, westward in the United States to the Turtle Mountains of North Dakota, the Black Hills of South Dakota, western Nebraska, central Kansas and Oklahoma, and the valley of the upper Colorado River (Fort Chadbourne, Coke County), Texas; very common northward, less abundant and of smaller size southward; abundant on the banks of streams flowing through the midcontinental plateau.

Largely planted as an ornamental and shade tree in the northern states, and rarely in western and northern Europe.

Ulmus ThomasiiSarg.

Fig. 283

Leavesobovate to oblong-oval, rather abruptly narrowed at apex into a short broad point, equally or somewhat unequally rounded, cuneate or subcordate at base, and coarsely doubly serrate, when they unfold pilose on the upper surface and covered on the lower with soft white hairs, at maturity 2′—2½′ long, ¾′—1′ wide, thick and firm, smooth, dark green and lustrous above, paler and soft-pubescent below, especially on the stout midrib and the numerous straight veins running to the point of the teeth and connected by obscure cross veinlets; turning in the autumn bright clear yellow; petioles pubescent, about ¼′ in length; stipules ovate-lanceolate, conspicuously veined, light green, marked with dark red on the margins above the middle, ⅔′ long, clasping the stem by their abruptly enlarged cordate base conspicuously dentate with 1—3 prominent teeth on each side, falling when the leaves are half grown.Flowerson elongated slender drooping pedicels often ½′ long, in 2—4, usually in 3-flowered, puberulous cymes becoming more or less racemose by the lengthening of the axis of the inflorescence, and when fully grown sometimes 2′ in length; calyx green, divided nearly to the middle into 7 or 8 rounded dark red scarious lobes; anthers dark purple; ovary coated with long pale hairs most abundant on the margins; styles light green.Fruitripening when the leaves are about half grown, ovoid or obovoid-oblong, ½′ long, with a shallow open notch at the apex, obscurely veined, pale pubescent, ciliate on the slightly thickened border of the broad wing, the margin of the seminal cavity scarcely thickened.

A tree, 80°—100° high, with a trunk occasionally 3° in diameter, and often free of branches for 60°, short stout spreading branches forming a narrow round-topped head, and slender rigid branchlets, light brown when they first appear, and coated with soft pale pubescence often persistent until their second season, becoming light reddish brown, puberulous or glabrous and lustrous in their first winter, and marked by scattered oblong lenticels and large orbicular or semiorbicular leaf-scars displaying an irregular row of 4—6 fibro-vascular bundle-scars, ultimately dark brown or ashy gray, and usually furnished with 3 or 4 thick corky irregular wings often ½′ broad, and beginning to appear in their first or more often during their second year.Winter-budsovoid, acute, ¼′ long, with broadly ovate rounded chestnut-brown scales pilose on the outer surface, ciliate on the margins, the inner scales becoming ovate-oblong to lanceolate, and ½′ long, often dentate at the base, with 1 or 2 minute teeth on each side, bright green below the middle, marked with a red blotch above, and white and scarious at the apex.Bark¾′—1′ thick, gray tinged with red, and deeplydivided by wide irregular interrupted fissures into broad flat ridges broken on the surface into large irregularly shaped scales.Woodheavy, hard, very strong and tough, close-grained, light clear brown often tinged with red, with thick lighter colored sapwood; largely employed in the manufacture of many agricultural implements, for the framework of chairs, hubs of wheels, railway-ties, the sills of buildings, and other purposes demanding toughness, solidity and flexibility.

Distribution.Dry gravelly uplands, low heavy clay soils, rocky slopes and river cliffs; Province of Quebec westward through Ontario, the southern peninsula of Michigan and central Wisconsin to northeastern Nebraska, western Missouri and eastern Kansas, and southward to northern New Hampshire, southern Vermont, western New York, (valley of the Genesee River), northern New Jersey, southern Ohio (near Columbus, Franklin County), and central Indiana; rare in the east and toward the extreme western and southern limits of its range.

Occasionally planted as a shade and ornamental tree in the northern states.

Fig. 284

Leavesovate-oblong to oblong-lanceolate, often somewhat falcate, acute or acuminate, unequally cuneate or rounded or subcordate at base, and coarsely doubly serrate with incurved teeth, when they unfold pale green often tinged with red, coated on the lower surface with soft white pubescence and glabrous or nearly so on the upper surface, at maturity thick and firm or subcoriaceous, dark green and smooth above, pale and soft-pubescent below, especially on the stout yellow midrib and numerous straight prominent veins often forked near the margins of the leaf and connected by rather conspicuous reticulate veinlets; turning yellow in the autumn; their petioles stout, pubescent, ⅓′ in length; stipules linear-obovate, thin and scarious, tinged with red above the middle, often nearly 1′ long.Flowerson drooping pedicels, in short few-flowered fascicles; calyx glabrous and divided nearly to the middle into 5 broad ovate rounded lobes as long as the hoary-tomentose ovary raised on a short slender stipe.Fruitripening before or with the unfolding of the leaves, oblong, ⅓′ in length, contracted at base into a long slender stalk, gradually narrowed and tipped at apex with long incurved awns, and covered with long white hairs most numerous on the thickened margin of the narrow wing;seedovoid, pointed, ⅛′ long, pale, chestnut-brown, slightly thickened into a narrow wing-like margin.

A tree, occasionally 80°—100° but usually not more than 40°—50° high, with a trunk 2°—3° in diameter, short stout straight or erect branches forming a narrow oblong rather openround-topped head, and slender branchlets glabrous or puberulous and light green tinged with red when they first appear, becoming light reddish brown or ashy gray and glabrous, or on vigorous individuals frequently pilose in their first winter, marked by occasional small orange-colored lenticels and by small elevated horizontal semiorbicular leaf-scars, sometimes naked, more often furnished with usually 2 thin corky wings beginning to grow during their first or more often during their second season, abruptly arrested at the nodes, often ½′ wide, and persistent for many years.Winter-budsslender, acute, ⅛′ long, dark chestnut-brown, with glabrous or puberulous scales, those of the inner ranks becoming oblong or obovate, rounded and tipped with a minute mucro, thin and scarious, light red, especially above the middle, and ½′ long.Barkrarely exceeding ¼′ in thickness, light brown tinged with red, and divided by irregular shallow fissures into flat ridges covered by small closely appressed scales.Woodheavy, hard, not strong, close-grained, difficult to split, light brown, with thick lighter colored sapwood; sometimes employed for the hubs of wheels and the handles of tools. Ropes used for fastening the covers of cotton bales are sometimes made from the inner bark.

Distribution.Usually on dry gravelly uplands, less commonly in alluvial soil on the borders of swamps and the banks of streams, and occasionally in inundated swamps; southeastern Virginia, southwestern Indiana, southern Illinois (Richland and Johnson Counties) and southern Missouri, and southward to central Florida (Lake County), and the valley of the Guadalupe River, Texas; ranging westward in Oklahoma to Garfield County (near Kingfisher,G. W. Stevens).

Often planted as a shade-tree in the streets of towns and villages of the southern states.

Fig. 285

Leavesovate-oblong, abruptly contracted into a long slender point, rounded at base on one side and short-oblique on the other, and coarsely doubly serrate with incurved callous-tipped teeth, when they unfold thin, coated below with pale pubescence, pilose above with scattered white hairs, at maturity thick and firm, dark green and rugose with crowded sharp-pointed tubercles pointing toward the apex of the leaf above, soft, smooth, and coated below, especially on the thin midrib and in the axils of the slender straight veins with white hairs, 5′—7′ long, 2′—3′ wide; turning a dull yellow color in the autumn; petioles stout, pubescent, ⅓′ in length; stipules obovate-oblong to oblong-lanceolate, thin and scarious, pale-pubescent, and tipped with clusters of rusty brown hairs.Flowerson short pedicels, in crowded fascicles; calyx green, covered with pale hairs, divided into 5—9short rounded thin equal lobes; stamens with slender light yellow slightly flattened filaments and dark red anthers; stigmas slightly exserted, reddish purple, papillose with soft white hairs.Fruitripening when the leaves are about half grown, semiorbicular, rounded and bearing the remnants of the styles or slightly emarginate at apex, rounded or cuneate at base, ½′ broad, the seminal cavity coated with thick rusty brown tomentum, the broad thin wing obscurely reticulate-veined, naked on the thickened margin, and marked by the dark conspicuous horizontal line of union of the two carpels;seedovoid, with a large oblique pale hilum, a light chestnut-brown coat produced into a thin border wider below than above the middle of the seed.

A tree, 60°—70° high, with a trunk occasionally 2° in diameter, spreading branches forming a broad open flat-topped head, and stout branchlets bright green, scabrate, and coated with soft pale pubescence when they first appear, becoming light brown by midsummer, often roughened by small pale lenticels, and in their first winter ashy gray, orange color or light red-brown, and marked by large elevated semiorbicular leaf-scars showing the ends of 3 conspicuous equidistant fibro-vascular bundles, ultimately dark gray or brown.Winter-budsovoid, obtuse, ¼′ long, with about 12 scales, the outer broadly ovate, rounded, dark chestnut-brown, and covered by long scattered rusty hairs, the inner when fully grown ½′ long, ⅛′—¼′ wide, light green, strap-shaped, rounded and tipped at the apex with tufts of rusty hairs, puberulous on the outer surface, slightly ciliate on the margins, gradually growing narrower and passing into the stipules of the upper leaves.Barkfrequently 1′ thick, dark brown tinged with red, divided by shallow fissures and covered by large thick appressed scales.Woodheavy, hard, strong, very close-grained, durable, easy to split, dark brown or red, with thin lighter colored sapwood; largely used for fence-posts, railway-ties, the sills of buildings, the hubs of wheels, and in agricultural implements. The thick fragrant inner bark is mucilaginous and demulcent, and is employed in the treatment of acute febrile and inflammatory affections.

Distribution.Banks of streams and low rocky hillsides in deep rich soil; comparatively common in the valley of the St. Lawrence River, Province of Quebec, and through Ontario to northern and eastern South Dakota, northeastern and eastern Nebraska, southeastern Kansas, and Oklahoma to the valley of the Canadian River (McClain County), and southward to western Florida, central Alabama and Mississippi, western Louisiana and the valley of the upper Guadalupe (Kerr County) and Leon Rivers (Comal County), Texas; in the South Atlantic states not common and mostly confined to the middle districts, ascending to altitudes of 2000° on the southern Appalachian foothills.

Fig. 286

Leaveselliptic to ovate, acute or rounded at apex, unequally rounded or cuneate and often oblique at base, coarsely and unequally doubly serrate with callous-tipped teeth, when they unfold thin, light green tinged with red, pilose above and covered below with soft pale pubescence, at maturity thick and subcoriaceous, dark green, lustrous and roughened by crowded minute sharp-pointed tubercles on the upper surface and soft pubescent on the lower surface, 1′—2′ long, ½′—1′ wide, with a stout yellow midrib, and prominent straight veins connected by conspicuous more or less reticulate cross veinlets; usually turning bright yellow late in the autumn; petioles stout, tomentose, ¼′—½′ in length; stipules ½′ long, linear-lanceolate, red and scarious above, clasping the stem by their green and hairy bases, deciduous when the leaves are about half grown.Flowersusually opening in August and sometimes also in October, on slender pedicels ⅓′—½′ long and covered with white hairs, in 3—5-flowered pedunculate fascicles; calyx divided to below the middle into oblong pointed lobes hairy at base; ovary hirsute, crowned with two short slightly exserted stigmas.Fruitripening in September and rarely also in November, oblong, gradually and often irregularly narrowed from the middle to the ends, short-stalked, deeply notched at apex, ⅓′ to nearly ½′ long, covered with soft white hairs, most abundant on the slightly thickened margin of the broad wing;seedoblique, pointed, and covered by a dark chestnut-brown coat.

A tree, often 80° high, with a tall straight trunk 2°—3° in diameter, sometimes free ofbranches for 30° or 40°, divided into numerous stout spreading limbs forming a broad inversely conic round-topped head of long pendulous branches, or while young or on dry uplands a compact round head of drooping branches, and slender branchlets, tinged with red and coated with soft pale pubescence when they first appear, becoming light reddish brown, puberulous and marked by scattered minute lenticels and by small elevated semiorbicular leaf-scars showing the ends of 3 small fibro-vascular bundles, and furnished with 2 corky wings covered with lustrous brown bark, about ¼′ broad and continuous except when abruptly interrupted by lateral branchlets, or often irregularly developed.Winter-budsbroadly ovoid, acute, ⅛′ long, with closely imbricated chestnut-brown scales slightly puberulous on the outer surface, those of the inner ranks at maturity oblong, concave, rounded at apex, thin, bright red, sometimes ¾′ long.Barksometimes nearly 1′ thick, light brown slightly tinged with red, and deeply divided by interrupted fissures into broad flat ridges broken on the surface into thick scales.Woodheavy, hard, strong, brittle, light brown tinged with red, with thick lighter colored sapwood; in central Texas used in the manufacture of the hubs of wheels, for furniture, and largely for fencing.

Distribution.Valley of the Sunflower River, Mississippi (Morehead, Sunflower County), through southern Arkansas, and Texas to Nuevo Leon, ranging in western Texas from the coast to the valley of the Pecos River; in Arkansas usually on river cliffs and low hillsides, and in Texas near streams in deep alluvial soil and on dry limestone hills; the common Elm-tree of Texas and of its largest size on the bottom-lands of the Guadalupe and Trinity Rivers.

Occasionally planted as a shade-tree in the streets of the cities and towns of Texas.

Fig. 287

Leavesoblong to oblong-obovate, acuminate, very oblique at base, coarsely and doubly crenulate-serrate, when they unfold coated below with shining white hairs and puberulous above, at maturity thin and firm in texture, yellow-green, glabrous and lustrous on the upper surface, pale and puberulous on the midrib and principal veins below, 2′—4′ long, 1′—1¾′ wide, with a prominent yellow midrib, about 20 pairs of primary veins extending obliquely to the points of the teeth and often forked near the margins of the leaf, and numerous reticular veinlets; turning clear orange-yellow in the autumn; petioles stout, about ¼′ in length; stipules abruptly narrowed from broad clasping bases, linear-lanceolate, usually about ¼′ long, persistent until the leaves are nearly fully grown.Flowersopening inSeptember on slender conspicuously jointed pedicels often ⅛′ long, in many-flowered glabrous racemes from 1′—1½′ in length; calyx 6-parted to the base, with oblong-obovate red-brown divisions rounded at apex; ovary sessile, narrowed below, villose.Fruitripening early in November, stipitate, oblong-elliptic, deeply divided at apex, fringed on the margins with long silvery white hairs, about ½′ long.

A tree, 50°—60° high, with a trunk 2°—3° in diameter, comparatively small spreading or pendulous branches often forming a broad handsome head, and slender pendulous branchlets glabrous or occasionally puberulous when they first appear, brown, lustrous, and marked by occasional oblong white lenticels during their first year, becoming darker the following season and ultimately dark gray-brown, and often furnished with 2 or 3 thick corky wings developed during their second or third years.Winter-budsovoid, acute, ¼′ long, their outer scales oblong-obovate, dark chestnut-brown, glabrous, the inner often scarious on the margins, pale yellow-green, lustrous and sometimes ¾′ long when fully grown.Bark¼′—⅜′ thick, light brown slightly tinged with red, and divided by shallow fissures into broad flat ridges broken on the surface into large thin closely appressed scales.Woodhard, close-grained, very strong and tough, light red-brown, with pale yellow sapwood.

Distribution.Limestone hills and river banks; rare and local; eastern (near Pikeville, Pike County) and southern Kentucky (Bowling Green, Warren County); banks of the Cumberland River, near Clarksville and Nashville, Tennessee; northeastern Georgia (cliffs of the Coosa River, near Rome, Floyd County); northern Alabama (Madison, Jefferson and Tuscaloosa Counties); valley of the Arkansas River (near Van Buren, Crawford County,G. M. Brown) and northwestern Arkansas (Sulphur Springs, Benton County, and Boston Mountains near Jasper, Newton County,E. J. Palmer); eastern Oklahoma (near Muskogee, Muskogee County,B. H. Slavin); southwestern (Grand Tower, Jackson County,H. A. Gleason) and southern Illinois (Richland County,R. Ridgway).

Occasionally planted as a shade-tree in the streets of cities in northern Georgia and northern Alabama; hardy in Eastern Massachusetts.

A tree, with scaly puberulous branchlets roughened by scattered pale lenticels, and at the end of their first season by small nearly orbicular leaf-scars marked by a row of fibro-vascular bundle-scars, minute subglobose winter-buds covered by numerous thinclosely imbricated chestnut-brown scales, the outer more or less scarious on the margins, the inner accrescent, becoming at maturity ovate-oblong, scarious, bright red, ⅓′—½′ long, marking in falling the base of the branchlet with pale ring-like scars. Leaves alternate, 2-ranked, ovate-oblong, acute or rounded at the narrowed apex, unequally cuneate or rounded at base, coarsely crenately serrate with unequal gland-tipped teeth, with numerous straight conspicuous veins forked near the margin and connected by cross reticulate veinlets more conspicuous below than above, when they unfold puberulous on the lower and pilose on the upper surface, at maturity thick or subcoriaceous and scabrate; petiolate with slender terete puberulous petioles; stipules lateral, free, ovate, scarious, bright red. Flowers polygamo-monœcious, the staminate fascicled in the axils of the outer scales of leaf-bearing buds, short-pedicellate, the pistillate or perfect on elongated puberulous pedicels in the axils of the leaves of the year in 1—3-flowered fascicles; pedicels without bracts; calyx campanulate, divided nearly to the base into 4 or 5 lobes rounded at apex, greenish yellow often tinged with red; stamens inserted under the ovary in the pistillate flower, sometimes few or 0; filaments filiform, erect, exserted; anthers broadly ovate, emarginate, cordate; ovary ovoid, stipitate, glandular-tuberculate, narrowed into a short style divided into 2 elongated reflexed stigmas papillo-stigmatic on the inner face, 0 in the staminate flower; ovule anatropous; micropyle extrorse, superior. Fruit an oblong oblique drupe, narrowed below into a short stipe, inclosed at the base by the withered calyx, crowned by the remnants of the style, its pericarp crustaceous, prominently ribbed on the anterior and posterior faces, irregularly tuberculate with elongated projections, and light chestnut-brown; seed ovoid, oblique, pointed at apex, rounded below, without albumen; testa thin, lustrous, dark brown or nearly black, of two coats; raphe inconspicuous; embryo erect; cotyledons thick, unequal, bright orange color, the apex of the larger hooded and slightly infolding the smaller, much longer than the minute radicle turned toward the linear pale hilum.

The genus is represented by a single species.

The generic name is in memory of Johann Jacob Planer, a German botanist and physician of the eighteenth century.

Fig. 288

Leaves2′—2½′ long, ¾′—1′ wide, on petioles varying from ⅛′—¼′ in length, dark dull green on the upper surface, paler on the lower surface, with a yellow midrib and veins.Flowersappearing with the leaves.Fruitripening in April, ⅓′ long.

A tree, 30°—40° high, with a short trunk rarely exceeding 20′ in diameter, rather slender spreading branches forming a low broad head, and branchlets brown tinged with red whenthey first appear, dark red in their first winter, and ultimately reddish brown or ashy gray.Barkabout ¼′ thick, light brown or gray, separating into large scales disclosing in falling the red-brown inner bark.Woodlight, soft, not strong, close-grained, light brown, with thick nearly white sapwood of 20—30 layers of annual growth.

Distribution.Swamps covered with water during several months of the year, or low river banks; valley of the Cape Fear River, North Carolina, southward to northern Florida (Bradford County) and westward usually not far from the coast through the Gulf states to the valleys of the Navasota (Brazos County) and of the Colorado (Matagorda County) Rivers, Texas, and northward through western Louisiana, eastern Oklahoma, and Arkansas to southeastern Missouri, northeastern Mississippi (near Iuka, Tishomingo County,T. G. Harbison), northern Kentucky (Henderson County), and the valley of the lower Wabash River, Illinois; comparatively rare and confined to the coast plain in the Atlantic states; abundant and of its largest size in western Louisiana and southern Arkansas.

Trees or shrubs, with thin, smooth often more or less muricate bark, unarmed or spinose branchlets, and scaly buds. Leaves serrate or entire, 3-nerved in one species, membranaceous or subcoriaceous, deciduous; stipules lateral, free, usually scarious, inclosing their leaf in the bud, caducous. Flowers polygamo-monœcious or rarely monœcious, appearing soon after the unfolding of the leaves, minute, pedicellate, on branches of the year, the staminate cymose or fascicled at their base, the pistillate solitary or in few-flowered fascicles from the axils of upper leaves; calyx divided nearly to the base into 4 or 5 lobes, greenish yellow, deciduous; stamens inserted on the margin of the discoid torus; filaments subulate, incurved in the bud, those of the sterile flower straightening themselves abruptly and becoming erect and exserted, shorter and remaining incurved in the perfect flower; anthers ovoid, attached on the back just above the emarginate base; ovary ovoid, sessile, green and lustrous, crowned with a short sessile style divided into diverging elongated reflexed acuminate entire lobes papillo-stigmatic on the inner face and mature before the anthers of the sterile flower, deciduous; minute and rudimentary in the staminate flower; ovule anatropous. Fruit an ovoid or globose drupe tipped with the remnants of the style, with thin flesh covered by a thick firm skin, and a thick-walled bony nutlet, reticulate-pitted in the American species. Seed filling the seminal cavity; albumen scanty, gelatinous, nearly inclosed between the folds of the cotyledons, or 0; testa membranaceous, of 2 confluent coats; chalaza colored, close to the minute hilum; embryo curved; cotyledons broad, foliaceous, conduplicate or rarely flat, variously folded, corrugate, incumbent, or inclosing the short superior ascending radicle.

Celtis is widely distributed through the temperate and tropical regions of the world, fifty or sixty species being distinguished.

Trees of the American species are often disfigured by gall-making insects which distort the buds and cause the production of dark broom-like clusters of short slender branchlets at the end of the branches.

Celtiswas the classical name of a species of Lotus.

Fig. 289

Leavesovate, short-acuminate or acute at apex, obliquely rounded at base, sharply serrate often only above the middle, thin, slightly pubescent below on the slender midrib and veins early in the season, becoming glabrous or nearly glabrous, 2½′—3½′ long, 1½′—2′ wide; turning yellow late in the autumn; petioles slender, glabrous, ⅓′—½′ in length.Flowerson drooping pedicels; calyx divided usually into 5 linear acute thin and scarious lobes rounded on the back, more or less laciniately cut, and often furnished with a tuft of pale hairs at apex; torus hoary-tomentose.Fruiton stems ½′—¾′ long, ripening in September and October and often remaining on the branches during the winter, subglobose, ovoid or obovoid, dark purple, ⅓′ in diameter, with a thick tough skin, dark orange-colored flesh and a thick-walled oblong pointed light brown slightly rugose nutlet;seedpale brown.

A tree, rarely more than 40°—50° high with a trunk usually not more than 2° in diameter, spreading often pendulous branches forming a round-topped head, and slender ridged light brown glabrous branchlets marked by oblong pale lenticels, and by horizontal semioval or oblong leaf-scars showing the ends of three fibro-vascular bundles, becoming darker and in their second or third year often dark red-brown.Winter-budsovoid, pointed, flattened, about ¼′ long, with three pairs of chestnut-brown ovate acute pubescent caducous scales closely imbricated in two ranks, increasing in size from without inward.Bark1′—1½′thick, smooth, dark brown, and more or less thickly covered and roughened by irregular, wart-like excrescences or by long ridges also found on the large branches.Woodheavy, rather soft, not strong, coarse-grained, clear light yellow, with thick lighter-colored sapwood; used for fencing and in the manufacture of cheap furniture.

Distribution.Rocky hills and ridges; New England (rare) to Virginia and westward to Iowa, eastern North Dakota, southwestern Missouri and northwestern Kansas.

Often planted in some of its forms as a shade and ornamental tree in the towns of the Mississippi valley and occasionally in the eastern states and in Europe.

Well distinguished by its large dark fruit,Celtis occidentalisis so variable in the shape of its leaves that two principal varieties are described as follows:

Celtis caninaRaf.

Fig. 290

Leavesoblong-ovate, gradually narrowed into a long acuminate point, obliquely rounded or unsymmetrically cuneate at base, finely serrate, glabrous or rarely pilose along the midrib and veins below, 2½′—6′ long and ¾′—2½′ wide; petioles slender, glabrous or rarely pubescent, ½′—¾′ long.

A tree, often 80°—100° high; more common than the other forms ofCeltis occidentalis.

Distribution.Rich wooded slopes and bottoms, or eastward on rocky ridges; Province of Quebec to eastern Nebraska, and southward to the coast of Massachusetts, western New York, southern Ohio, southern Indiana and Illinois, southwestern Missouri, southwestern Oklahoma (Snyder, Kiowa County), and in northwestern Georgia.

Celtis crassifoliaLam.

Fig. 291

Leavesthicker, long-acuminate, obliquely rounded at base, usually more coarsely serrate, rarely nearly entire, rough on the upper surface, pilose below along the prominent midrib and veins, 3½′—5′ long, 2′—2½′ wide, much smaller in the Rocky Mountain region; petioles villose-pubescent, rarely glabrous, ¼′—½′ in length, much shorter than the pubescent pedicels of the fruit.

A tree, 100°—120° high; with pubescent or glabrous branchlets; rarely shrubby. The most widely distributed form ofCeltis occidentalis.

Distribution.Wooded slopes and rich bottoms; Virginia and along the Appalachian Mountains to North Carolina and westward to southern Minnesota, Missouri, centralKansas, eastern and northwestern Oklahoma, central Nebraska, North and South Dakota, cañons of the Big Horn Mountains, Wyoming, and northwestern Idaho, and southward to Dallas County, Alabama, and eastern Texas.

Often cultivated in towns of the Mississippi Valley and in western Europe, and occasionally in the eastern states.

Celtis rugulosaRydb.

Fig. 292

Leavesbroadly ovate to oblong-ovate, acuminate, obliquely rounded or unsymmetrically subcordate at base, coarsely serrate, rough on the upper surface, pale and covered below with a network of reticulate veinlets inconspicuous early in the season, later becoming prominent, glabrous or sparingly pilose along the under side of the stout midrib and primaryveins, 2′—2½′ long, 1′—2′ wide; petioles stout, slightly pubescent, ¼′—½′ in length.Flowerson slender pubescent pedicels; calyx divided into five linear acute scarious lobes laciniately cut at apex; torus hoary-tomentose.Fruiton slender drooping slightly pubescent or glabrous pedicels, ⅓′—½′ in length, subglobose to ellipsoid, light orange-brown, lustrous, ⅓′ in diameter.

A small tree or shrub rarely more than 20′ high, with slender slightly pubescent or glabrous red-brown branchlets marked by small pale lenticels, becoming ashy gray in their second or third year.Barkrough, red-brown or gray.

Distribution.Dry hillsides and rocky river banks; eastern Oregon from the valley of the Deschutes and Columbia Rivers to the cañon of Snake River, Whitman County, Washington, and to Big Willow Creek, Cañon County, western Idaho; on the western foothills of the Wasatch Mountains, in the cañon of Grand River, and in Diamond Valley, Utah; southern California, near Independence, Inyo County, Hackberry Cañon, Kern County, and Things Valley at base of Laguna Mountain, near Campo, southern San Diego County; on Cedros Island, and in northern Lower California; rim of the Grand Cañon, Arizona, and on the eastern foothills of the Rocky Mountains of Colorado.

Occasionally planted in the towns of western Washington, and when cultivated said to grow in good soil into a larger and more shapely tree with thinner leaves.

Celtis HelleriSmall.

Fig. 293

Leavesoblong-ovate, acuminate or acute, cordate or obliquely cordate or rounded at base, entire, or crenately serrate on vigorous shoots, rough above, pale and clothed below with white hairs, becoming by midsummer thick and covered below with a conspicuous network of reticulate veinlets, 1½′—3′ long, ¾′—2′ wide; petioles densely villose-pubescent, ¼′—½′ in length.Flowersopening toward the end of March on pubescent pedicels; calyx divided into five oblong scarious lobes narrowed and rounded at apex; torus tomentose.Fruiton slender tomentose stems ¼′—⅔′ long, ripening in September and persistent on the branches until spring, subglobose to ellipsoid, dark reddish brown, lustrous, ¼′ in diameter.

A tree, occasionally 30° high, with a trunk rarely more than 12′—18′ in diameter, stout spreading branches forming a broad open irregular head, and slender pubescent branchlets roughened by numerous small lenticels, becoming darker and glabrous in their second season.Barkof the trunk and large branches dark and covered with high thick wart-like excrescences and ridges.Woodnot strong nor durable, of little value even for fuel.

Distribution.Rich bottom-lands and on low adjacent hills of streams flowing southward from the Edward’s Plateau (Goliad, San Antonio, New Braunfels, San Marcos) and near Austin, Travis County, Texas.

Fig. 294

Leavesbroadly ovate, acute or acuminate, obliquely rounded at base, entire, thick, dark green and rough or rarely smooth on the upper surface, yellow-green and conspicuously reticulate-venulose and sparingly pilose along the prominent midrib and veins on the lower surface, 1¼′—3′ long, ¾′—1½′ wide; petioles stout, ⅛′—¼′ in length, more or less densely pubescent.Flowersnot seen.Fruiton pubescent pedicels ⅓′—½′ in length, ripening in September, subglobose to ellipsoid, orange-red or yellow, lustrous, ¼′ in diameter.

A tree, rarely 30° high with stout ascending branches forming an open irregular head, and slender red-brown branchlets tomentose or pubescent early in their first season and pubescent or glabrous in their second year; or often a shrub.Barkthick and rough.

Distribution.Dry limestone hillsides, rocky ridges and cañon slopes, western Texas, from the valley of the upper Rio Frio, Uvalde County, to Oklahoma (Ozark region, near Page, Le Flore County to the southwestern borders of the state); in mountain ravines through southern New Mexico, and in southern central and northeastern Arizona.

A variety with more pubescent serrate leaves, those on vigorous shoots mostly cordate at base and covered above with short white hairs, is distinguished as var.vestitaSarg. A small tree with slender pubescent branchlets and a trunk 12′—15′ in diameter. In low ground, along the North Fork of the Canadian River, near Canton, Blaine County, Oklahoma.

Celtis mississippiensisSpach.

Fig. 295

Leavesoblong-lanceolate, long-pointed and acuminate at apex, unsymmetrically rounded or cuneate or obliquely cuneate at base, often falcate, entire or furnished with a few teeth near the apex or serrate (var.SmalliiSarg.), thin, smooth, glabrous or rarely rough above, light green on both surfaces, 2½′—5′ long and ¾′—1½′ wide, with a narrow yellowmidrib, slender veins arcuate and united near the margins, and inconspicuous reticulate veinlets; petioles slender, glabrous, ¼′—½′ in length.Flowerson slender glabrous pedicels; calyx divided into five ovate-lanceolate glabrous or puberulous scarious lobes furnished at apex with tufts of long white hairs.Fruiton glabrous pedicels shorter or slightly longer than the petioles, ripening in September, short-oblong to ellipsoid or obovoid, orange-red or yellow, ¼′ in diameter; nutlet slightly rugose.

A tree, 60°—80° high, with a trunk 2°—3° in diameter, spreading or pendulous branches forming a broad head, and slender branchlets light green, glabrous or pubescent when they first appear, and during their first winter bright reddish brown, rather lustrous and marked by oblong pale lenticels and narrow elevated horizontal leaf-scars showing the ends of three fibro-vascular bundles; often much smaller.Winter-budsovoid, pointed,1/16′—⅛′ long, with chestnut-brown puberulous scales.Bark½′—⅔′ thick, pale gray and covered with prominent excrescences.Woodsoft, not strong, close-grained, light yellow, with thick lighter-colored sapwood; commercially confounded with the wood ofCeltis occidentalisand its varieties, and used for the same purposes.

Distribution.Coast of Virginia to the Everglades Keys of southern Florida, through the Gulf states to the valley of the lower Rio Grande in Nuevo Leon, and through eastern Texas, Arkansas and Missouri to eastern Oklahoma to the valley of the Washita River (Zarvin County) and to Kiowa County, eastern Kansas, central Tennessee and Kentucky, and to southern Illinois and Indiana; in Bermuda.

Often planted as a shade and street tree in the valley of the Mississippi River and in Texas.

An arborescent form from the rocky banks of the Nueces River, western Texas, with shorter and thicker leaves is distinguished as var.brachyphyllaSarg.; and a small shrubby form with oblong-ovate cordate leaves and dark purplish fruit covered with a glaucous bloom, growing in deep sand in Callihan County, Texas, has been described as var.anomalaSarg. An Arizona form is

Celtis brevipesS. Wats.


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