BOVINÆ—CATTLE.
BOVINÆ—CATTLE.
These comprise the oxen, and wind up the hollow-horned ruminants as far as India is concerned. There are in the New World some other very interesting animals of this group, such as the musk-ox (Ovibos), and the prong-horned antelope (Antilocapra), which last so far resembles the Cervidæ that the horns, which are bifurcate, are also annually shed. They come off the bony core, on which the new horn is already beginning to form.
The Bovines are animals of large size, horned in both sexes, a very large and broad moist muffle, massive bodies and stout legs. The horns, which are laterally wide spread, are supported on cores of cellular bone, and are cylindrical or depressed at the base. The nose broad, with the nostrils at the side. The skull has no sub-orbital pit or fissure, and the bony orbit is prominent; grinders with a well-developed supplementary lobe; cannon bone short. In India, the groups into which this sub-family may be divided, are oxen, the buffaloes, and the yaks. There are no true bison in our limits, the commonly so-called bison being properly a wild ox. The taurine or Ox group is divided into theZebus, or humped domestic cattle;Taurus, humpless cattle with cylindrical horns; andGavæus, humpless cattle with flattened horns.
According to Dr. Jerdon, in some parts of India small herds of zebus have run wild. He says:—
"Localities are recorded in Mysore, Oude, Rohilkund, Shahabad, &c., and I have lately seen and shot one in the Doab near Mozuffernugger. These, however, have only been wild for a few years. Near Nellore, in the Carnatic, on the sea-coast there is a herd of cattle that have been wild for many years. The country they frequent is much covered with jungle and intersected with salt-water creeks and back-waters, and the cattle are as wild and wary as the most feral species. Their horns were very long and upright, and they were of large size. I shot one there in 1843, but had great difficulty in stalking it, and had to follow it across one or two creeks."
GENUS GAVÆUS.
GENUS GAVÆUS.
Massive head with large concave frontals, surmounted inG. gaurusby a ridge or crest of bone; horns flattened on the outer surface, corrugated at the base, and smooth for the rest of the two-thirds, or a little more; wide-spreading and recurved at the tips, forming a crescent; greenish grey for the basal half, darker towards the tips, which are black; muffle small; dewlap small or absent; the spinous processes of the dorsal vertebræ are greatly developed down to about half the length of the back; legs small under the knee, and white in colour; hoofs small and pointed, leaving a deer-like print in the soil, very different to the splay foot of the buffalo.
NO. 464. GAVÆUS GAURUS.The Gaur, popularly called Bison(Jerdon's No. 238).
NO. 464. GAVÆUS GAURUS.The Gaur, popularly called Bison(Jerdon's No. 238).
NATIVENAMES.—GaororGaori-gai,Bun-boda, Hindi;BodaandBunparrain the Seonee and Mandla districts;Pera-maooof Southern Gonds;Gaoiya, Mahrathi;Karkona, Canarese;Katuyeni, Tamil;Jangli-kulghain Southern India;Pyoungin Burmah;Salandangin the Malay countries. Horsfield gives the following names under hisBibos asseel:As'l Gayal, Hindi;Seloi, Kuki;P'hanjof the Mughs and Burmese, and some others which he considers doubtful.
HABITAT.—Regarding this, I quote at length from Jerdon, whose inquiries were carefully made. He says: "The gaur is an inhabitant of all the large forests of India, from near Cape Comorin to the foot of the Himalayas. On the west coast of India it is abundant all along the Syhadr range on Western Ghâts, both in the forests at the foot of the hills, but more especially in the upland forests and the wooded country beyond the crest of the Ghâts. The Animally hills, the Neilgherries, Wynaad, Coorg, the Bababooden hills, the Mahableshwar hills, are all favourite haunts of this fine animal. North of this, it occurs to my own knowledge in the jungles on the Taptee river and the neighbourhood, and north of the Nerbudda; a few on the deeper recesses of the Vindhian mountains. On the eastern side of the peninsula it is found in the Pulney and Dindigul hills, the Shandamungalum range, the Shervaroys, and some of the hill ranges near Vellore and the borders of Mysore. North of this, the forest being too scanty, it does not occur till the Kistna and Godavery rivers; and hence it is to be found in suitable spots all along the range of Eastern Ghâts to near Cuttack and Midnapore, extending west far into Central India, and northwards towards the edge of the great plateau which terminates south of the Gangetic valley. According to Hodgson it also occurs in the Himalayan Terai, probably however only towards the eastern portion, and here it is rare, for I have spoken to many sportsmen who have hunted in various parts of the Terai, from Sikhim to Rohilkund, and none have ever come across the gaur at the foot of the Himalayas."—'Mam. of Ind.,' p. 303. (See alsoAppendix C.)
In the Central provinces the gaur is found in several parts of the bamboo-clad spurs of the Satpura range. My experience of the animal is limited to the Seonee district, where it is restricted to the now closely preserved forests of Sonawani in the south-east bend of the range, and a few are to be seen across occasionally, near the old fort of Amodagarh, on the Hirri river.
It is also more abundant on the Pachmari and Mahadeo hills. On the east of the Bay of Bengal it is found from Chittagong through Burmah to the Malayan peninsula. It was considered that the gaur of the eastern countries was a distinct species, and is so noted in Horsfield's Catalogue, and described at some length under the name ofBibos asseel; but it appears that all this distinction was founded on the single skull of a female gaur, and is an instance of the proneness of naturalists to create new species on insufficient data. He himself remarks that when the skin was removed it was evident that the animal was nearly related toGavæus gaurus, or, as he calls it,Bibos cavifrons. Mr. G. P. Sanderson shot a fine old male of what he supposed to be the wildgayal, and he says: "I can state that there was not one single point of difference in appearance or size between it and the bison of Southern India, except that the horns were somewhat smaller than what would have been looked for in a bull of its age in Southern India;" and this point was doubtless an individual peculiarity, for Blyth, in his 'Catalogue of the Mammals of Burmah,' says: "Nowhere does this grand species attain a finer development than in Burmah, and the horns are mostly short and thick, and very massive as compared with those of the Indian gaurs, though the distinction is not constant on either side of the Bay of Bengal."
Jerdon supposes it to have existed in Ceylon till within the present century, but I do not know on what data he founds his assertion.
DESCRIPTION.—I cannot improve on Jerdon's description, taken as it is from the writings of Hodgson, Elliot, and Fisher, so I give it as it stands, adding a few observations of my own on points not alluded to by them:—
"The skull is massive; the frontals large, deeply concave, surmounted by a large semi-cylindric crest rising above the base of the horns. There are thirteen pairs of ribs.[40]The head is square, proportionately smaller than in the ox; the bony frontal ridge is five inches above the frontal plane; the muzzle is large and full, the eyes small, with a full pupil (? iris) of a pale blue colour. The whole of the head in front of the eyes is covered with a coat of close short hair, of a light greyish-brown colour, which below the eyes is darker, approaching almost to black; the muzzle is greyish and the hair is thick and short; the ears are broad and fan-shaped; the neck is sunk between the head and back, is short, thick, and heavy. Behind the neck and immediately above the shoulder rises a gibbosity or hump of the same height as the dorsal ridge. This ridge rises gradually as it goes back, and terminates suddenly about the middle of the back; the chest is broad; the shoulder deep and muscular; the fore-legs short, with the joints very short and strong, and the arm exceedingly large and muscular; the hair on the neck and breast and beneath is longer than on the body, and the skin of the throat is somewhat loose, giving the appearance of a slight dewlap; the fore-legs have a rufous tint behind and laterally above the white. The hind-quarters are lighter and lower than the fore, falling suddenly from the termination of the dorsal ridge; the skin of the neck, shoulders, and thigh is very thick, being about two inches and more.
"The cow differs from the bull in having a slighter and more graceful head, a slender neck, no hump; and the points of the horns do not turn towards each other at the tip, but bend slightly backwards, and they are much smaller; the legs too are of a purer white. The very young bull has the forehead narrower than the cow, and the bony frontal ridge scarcely perceptible. The horns too turn more upwards. In old individuals the hair on the upper parts is often worn off. The skin of the under parts when uncovered is deep ochrey-yellow."—'Mammals of India,' p. 302.
40 The true bison has fourteen pairs of ribs.—R. A. S.
The fineness of the leg below the knee is another noticeable feature, and also the well-formed pointed hoof, which leaves an imprint like that of a large deer. Mr. Sanderson states in his book that the bison, after a sharp hunt, gives out an oily sweat, and in this peculiarity he says it differs from domestic cattle, which never sweat under any exertion. This I have not noticed.
The period of gestation seems to be about the same as that of the domestic cow, and the greatest number of calves are born in the summer.
SIZE.—I cannot speak personally, for I regret now that I took no measurements in the days when I was acquainted with these magnificent animals, but the experiences of others I give as follows:—
Sir Walter Elliot gives—
I give the measurements of two fine heads:—
The following careful measurements are recorded by Mr. Blyth ('J. A. S. B.,' vol. xi., 1842, p. 588), and were furnished to him by Lieut. Tickell from the recently-killed animal, in order to assist in the setting up of the specimen in the Asiatic Museum:—
I feel tempted to let my pen run away with me into descriptions of the exciting scenes of the past in the chase of this splendid creature—the noblest quarry that the sportsman can have, and the one that calls forth all his cunning and endurance. As I lately remarked in another publication, I know of no other animal of which the quest calls forth the combined characteristics of the ibex, the stag and the tiger-hunter. Some of my own experiences I have described in 'Seonee;' but let those who wish to learn the poetry of the thing read the glowing, yet not less true pages of Colonel Walter Campbell's 'Old Forest Ranger;' and for clear practical information, combined also with graphic description, the works of Captain J. Forsyth and Mr. G. P. Sanderson ('The Highlands of Central India' and 'Thirteen years among the Wild Beasts').
The gaur prefers hilly ground, though it is sometimes found on low levels. It is extremely shy and retiring in its habits, and so quick of hearing that extreme care has to be taken in stalking to avoid treading on a dry leaf or stick. I know to my cost that the labour of hours may be thrown away by a moment of impatience. In spite of all the wondrous tales of its ferocity, it is as a rule a timid, inoffensive animal. Solitary bulls are sometimes dangerous if suddenly come upon. I once did so, and the bull turned and dashed up-hill before I could get a shot, whereas a friend of mine, to whom a similar thing occurred a few weeks before, was suddenly charged, and his gun-bearer was knocked over. The gaur seldom leaves its jungles, but I have known it do so on the borders of the Sonawani forest, in order to visit a small tank at Untra near Ashta, and the cultivation in the vicinity suffered accordingly.
Hitherto most attempts to rear this animal when young have failed. It is said not to live over the third year. Though I offered rewards for calves for my collection, I never succeeded in getting one. I have successfully reared most of the wild animals of the Central provinces, but had not a chance of trying the bison.
NO. 465. GAVÆUS FRONTALIS.The Mithun or Gayal.
NO. 465. GAVÆUS FRONTALIS.The Mithun or Gayal.
NATIVENAMES.—Gayal,GaviorGabi,Gabi-bichal(male),Gabi-gai(female);Bunerea-goruin Chittagong and Assam;Mithun.
HABITAT.—The hilly tracts east of the Brahmaputra, at the head of the Assam valley, the Mishmi hills, in hill Tipperah, Chittagong, and then southwards through Burmah to the hills bordering on the Koladyne river.
DESCRIPTION.—Very like the gaur at first sight, but more clumsy looking; similarly coloured, but with a small dewlap; the legs are white as in the last species. In the skull the forehead is not concave as in the gaur, but flat, and if anything rather convex. The back has a dorsal ridge similar to that of the gaur.
Thegayalis of a much milder disposition than thegaur, and is extensively domesticated, and on the frontiers of Assam is considered a valuable property by the people. The milk is rich and the flesh good. There are purely domesticatedmithunsbred in captivity, but according to many writers the herds are recruited from the wild animals, which are tempted either to interbreed, or are captured and tamed. In Dr. F. Buchanan Hamilton's MS. (seeHorsfield's 'Cat. Mammalia, E. I. C. Mus.') the following account is given: "These people (i.e. the inhabitants of the frontiers) have tame gayals, which occasionally breed, but the greater part of their stock is bred in the woods and caught; after which, being a mild animal, it is easily domesticated. The usual manner employed to catch the full-grown gayal is to surround a field of corn with a strong fence. One narrow entrance is left, in which is placed a rope with a running noose, which secures the gayal by the neck as he enters to eat the corn; of ten so caught perhaps three are hanged by the noose running too tight, and by the violence of their struggling. Young gayals are caught by leaving in the fence holes of a size sufficient to admit a calf, but which excludes the full-grown gayal; the calves enter by these holes, which are then shut by natives who are watching, and who secure the calves. The gayal usually goes in herds of from twenty to forty, and frequents dry valleys and the sides of hills covered with forest." Professor Garrod, in hisUngulatain Cassell's Natural History, quotes the following account from Mr. Macrae concerning the way in which the Kookies of the Chittagong hill regions catch the wild gayal: "On discovering a herd of wild gayals in the jungle they prepare a number of balls, the size of a man's head, composed of a particular kind of earth, salt and cotton. They then drive their tame gayals towards the wild ones, when the two herds soon meet and assimilate into one, the males of the one attaching themselves to the females of the other, andvice versâ. The Kookies now scatter their balls over such parts of the jungles as they think the herd most likely to pass, and watch its motions. The gayals, on meeting these balls as they pass along, are attracted by their appearance and smell, and begin to lick them with their tongues, and, relishing the taste of the salt and the particular earth composing them, they never quit the place till all the balls are consumed. The Kookies, having observed the gayals to have once tasted their balls, prepare a sufficient supply of them to answer the intended purpose, and as the gayals lick them up they throw down more; and it is to prevent their being so readily destroyed that the cotton is mixed with the earth and the salt. This process generally goes on for three changes of the moon or for a month and a-half, during which time the tame and the wild gayals are always together, licking the decoy balls, and the Kookie, after the first day or two of their being so, makes his appearance at such a distance as not to alarm the wild ones. By degrees he approaches nearer and nearer, until at length the sight of him has become so familiar that he can advance to stroke his tame gayals on the back and neck without frightening the wild ones. He next extends his hand to them and caresses them also, at the same time giving them plenty of his decoy balls to lick. Thus, in the short space of time mentioned, he is able to drive them, along with the tame ones, to hisparrahor village, without the least exertion of force; and so attached do the gayals become to theparrah, that when the Kookies migrate from one place to another, they always find it necessary to set fire to the huts they are about to abandon, lest the gayals should return to them from the new grounds."
NO. 466. GAVÆUSSONDAICUS.The Burmese Wild Ox.
NO. 466. GAVÆUSSONDAICUS.The Burmese Wild Ox.
NATIVENAME.—Tsoing, Burmese;Bantengof the Javanese.
HABITAT.—"Pegu, the Tenasserim provinces, and the Malayan peninsula, Sumatra, Borneo and Java; being domesticated in the island of Bali" (Blyth).
DESCRIPTION.—This animal resembles the gaur in many respects, and it is destitute of a dewlap, but the young and the females are bright chestnut. The bulls become black with age, excepting always the white stockings and a white patch on each buttock.
SIZE.—About the same as the last two species.
This animal has bred in captivity, and has also interbred with domestic cattle. Blyth says he saw in the Zoological Gardens of Amsterdam a bull, cow, and calf in fine condition. "The bull more especially has an indication of a hump, which, however, must be specially looked for to be noticed, and he has a broad and massive neck like the gaur, but no raised spinal ridge, nor has either of these species a deep dewlap like the gayal" ('Cat. Mamm. Burmah'). The banteng cow is much slighter in build, and has small horns that incline backwards, and she retains her bright chestnut colour permanently.
GENUS POEPHAGUS—THE YAK.
GENUS POEPHAGUS—THE YAK.
Somewhat smaller than the common ox, with large head; nose hairy, with a moderate sized bald muffle between nostrils; broad neck without dewlap; cylindrical horns; no hump or dorsal ridge, and long hair on certain parts of the body. Requires an intensely cold climate.
NO. 467. POEPHAGUS GRUNNIENS.The Yak or Grunting Ox.
NO. 467. POEPHAGUS GRUNNIENS.The Yak or Grunting Ox.
NATIVENAMES.—Yak,Bubul,Soora-goy,Dong, in Thibet;Bun-chowr, Hindi;Brong-dong, Thibetan.
HABITAT.—The high regions of Thibet and Ladakh, the valley of the Chang Chenmo, and the slopes of the Kara Koram mountains (Kinloch).
DESCRIPTION.—"In size it is somewhat less than the common or domestic ox. The head is large, and the neck proportionally broad, without any mane or dewlap, having a downward tendency; the horns are far apart, placed in front of the occipital ridge, cylindrical at the base, from which they rise obliquely outward and forward two-thirds of their length, when they bend inward with a semi-circular curve, the points being directed to each other from the opposite sides; the muffle is small; the border of the nostrils callous; the ears short and hairy. At the withers there is a slight elevation, but no protuberance or hump, as in the Indian ox. The dorsal ridge not prominent; body of full dimensions; rump and hinder parts proportionally large; limbs rather small and slender; hoofs smooth, square, and well defined, not expanded as in the musk-ox; anterior false hoofs small, posterior large; tail short, not reaching beyond the houghs, naked for some inches at the root, very bushy, lax, and expanded in the middle; colour black throughout, but varying in tint according to the character of the hairy covering; this, on the anterior parts, the neck, shoulders, back, and sides, is short, soft, and of a jet-black colour, but long, shaggy, pendulous, and shining on the sides of the anterior extremities, and from the medial part of the abdomen over the thighs to the hinder parts" (Horsfield, 'Cat. Mam. Ind. Mus.').
GENUS BUBALUS—THE BUFFALOS.
GENUS BUBALUS—THE BUFFALOS.
Horns very large, depressed and sub-trigonal at the base, attached to the highest line of the frontals, inclining upwards and backwards, conical towards the tip and bending upwards; muffle large, square. No hump or dorsal ridge; thirteen pairs of ribs; hoofs large.
NO. 468. BUBALUS ARNI.The Wild Buffalo(Jerdon's No. 239).
NO. 468. BUBALUS ARNI.The Wild Buffalo(Jerdon's No. 239).
NATIVENAMES.—Arna(male),Arni(female),Arna-bhàinsa,Jangli-bhains, Hindi;Mung, Bhagulpore;Gera-erumi, Gondi;Karboof the Malays;Moondingof the Sundanese.
HABITAT.—In the swampy terai at the foot of the hills from Oude to Bhotan, in the plains of Lower Bengal as far west as Tirhoot, in Assam and in Burmah, in Central India from Midnapore to Rajpore, and thence nearly to the Godavery; also in Ceylon.
DESCRIPTION.—This animal so closely resembles the common domesticated buffalo that it seems hardly necessary to attempt a description. The wild one may be a trifle larger, but every one in India is familiar with the huge, ungainly, stupid-looking creature, with its bulky frame, black and almost hairless body, back-sweeping horns, and long narrow head.
SIZE.—A large male will stand 19 hands at the shoulder and measure 10¼ feet from nose to root of tail, which is short, reaching only to the hocks. Horns vary greatly, but the following are measurements of large pairs: In the British Museum are a pair without the skull. These horns measure 6 feet 6 inches each, which would give, when on the head, an outer curve measurement of nearly 14 feet. Another pair in the British Museum measure on the skull 12 feet 2 inches from tip to tip and across the forehead, but these horns do not exactly correspond in length and shape.
The buffalo never ascends mountains like the bison, but keeps to low and swampy ground and open grass plains, living in large herds, which occasionally split up into smaller ones during the breeding season in autumn. The female produces one, or sometimes two in the summer, after a period of gestation of ten months.
Forsyth doubts their interbreeding with the domestic race, but I see no reason for this. The two are identically the same, and numerous instances have been known of the latter joining herds of their wild brethren; and I have known cases of the domestic animal absconding from a herd and running wild. Such a one was shot by a friend of mine in a jungle many miles from the haunts of men, but yet quite out of the range of the wild animal. Probably it had been driven from a herd. Domestic buffalo bulls are much used in the Central provinces for carrying purposes. I had them yearly whilst in camp, and noticed that one old bull lorded it over the others, who stood in great awe of him; at last one day there was a great uproar; three younger animals combined, and gave him such a thrashing that he never held up his head again. In a feral state he would doubtless have left the herd and become a solitary wanderer. Dr. Jerdon, in his 'Mammals of India,' says: "Mr. Blyth states it as his opinion that, except in the valley of the Ganges and Burrampooter, it has been introduced and become feral. With this view I cannot agree, and had Mr. Blyth seen the huge buffalos I saw on the Indrawutty river (in 1857), he would, I think, have changed his opinion. They have hitherto not been recorded, south of Raepore, but where I saw them is nearly 200 miles south. I doubt if they cross the Godavery river.
"I have seen them repeatedly, and killed several in the Purneah district. Here they frequent the immense tracts of long grass abounding in dense, swampy thickets, bristling with canes and wild roses; and in these spots, or in the long elephant-grass on the bank of jheels, the buffalos lie during the heat of the day. They feed chiefly at night or early in the morning, often making sad havoc in the fields, and retire in general before the sun is high. They are by no means shy (unless they have been much hunted), and even on an elephant, without which they could not be successfully hunted, may often be approached within good shooting distance. A wounded one will occasionally charge the elephant, and, as I have heard from many sportsmen, will sometimes overthrow the elephant. I have been charged by a small herd, but a shot or two as they are advancing will usually scatter them."
The buffalo is, I should say, a courageous animal—at least it shows itself so in the domesticated state. A number of them together will not hesitate to charge a tiger, for which purpose they are often used to drive a wounded tiger out of cover. A herdsman was once seized by a man-eater one afternoon a few hundred yards from my tent. His cows fled, but his buffalos, hearing his cries, rushed up and saved him.
The attachment evinced by these uncouth creatures to their keepers was once strongly brought to my notice in the Mutiny. In beating up the broken forces of a rebel Thakoor, whom we had defeated the previous day, I, with a few troopers, ran some of them to bay in a rocky ravine. Amongst them was a Brahmin who had a buffalo cow. This creature followed her master, who was with us as a prisoner, for the whole day, keeping at a distance from the troops, but within call of her owner's voice. When we made a short halt in the afternoon, the man offered to give us some milk; she came to his call at once, and we had a grateful draught, the more welcome as we had had nothing to eat since the previous night. That buffalo saved her master's life, for when in the evening the prisoners were brought up to court martial and sentenced to be hanged, extenuating circumstances were urged for our friend with the buffalo, and he was allowed to go, as I could testify he had not been found with arms in his hands; and I had the greatest pleasure in telling him to be off, and have nothing more to do with rebel Thakoors. Jerdon says the milk of the buffalo is richer than that of the cow. I doubt this. I know that in rearing wild animals buffalos' milk is better than cows' milk, which is far too rich, and requires plentiful dilution with water.
There is a very curious little animal allied to the buffalo, of which we have, or have had, a specimen in the Zoological Gardens at Alipore—theAnoa depressicornis; it comes from the Island of Celebes, and seems to link the buffalo with the deer. It is black, with short wavy hair.
Before passing on to the true Cervidæ I must here place an animal commonly called a deer, and generally classed as such—the musk-deer according to some naturalists. There is no reason, save an insufficient one, that this creature should be so called and classed, there being much evidence in favour of its alliance to the antelopes. In the first place it has a gall bladder, which the Cervidæ have not, with the exception, according to Dr. Crisp, of the axis ('P. Z. S.'). On the other hand it has large canine tusks like the muntjacs, deerlets, and water-deer, and, as these are all aberrant forms of the true Cervidæ, there is no reason why the same character should not be developed in the antelopes. Its hair is more of the goat than the deer, and the total absence of horns removes a decided proof in favour of one or the other. The feet are more like some of the Bovidæ than the generality of deer, with the exception, perhaps, ofRangifer(the reindeer), the toes being very much cloven and capable of grasping the rocky ground on which it is found. A very eminent authority, however, Professor Flower, is in favour of placing the musk-deer with the Cervidæ, and he instances the absence of horns as in favour of this opinion, for in none of the Bovidæ are the males hornless. There are many other points also, such as the fawns being spotted, some intestinal peculiarities, and the molar and premolar teeth being strictly cervine, which strengthen him in his opinion. (Seearticle on the structure and affinities of the musk-deer, 'P. Z. S.' 1879, p. 159.)
GENUS MOSCHUS—THE MUSK DEER.
GENUS MOSCHUS—THE MUSK DEER.
Canines in both sexes, very long and slender in the male; no horns; feet much cloven, with large false hoofs that touch the ground; the medium metacarpals fused into a solid cannon bone; in the skull the intermaxillaries join the nasals; hinder part of tarsus hairy; fur thick, elastic, and brittle; muffle large; no eye, feet, or groin-pits; a large gland or præputial bag under the stomach in the males, which contains the secretion known in commerce as "musk."
NO. 469. MOSCHUS MOSCHIFERUS.The Musk Deer.
NO. 469. MOSCHUS MOSCHIFERUS.The Musk Deer.
NATIVENAMES.—Kastura, Hindi;Rous,Roos, andKasturé, in Kashmir;La-lawa, Thibetan;Rib-jo, Ladakhi;Benain Kunawur (Jerdon);Mussuck-naba', Pahari (Kinloch).
HABITAT.—Throughout the Himalayas at elevations above 8000 feet, extending also through Central and Northern Asia as far as Siberia.
DESCRIPTION.—It is difficult to describe the colour of this animal, for it so constantly changes; and, as I do not know the creature personally, I think it better to give the recorded opinions of three writers who have had personal experience. Markham describes it as a dark speckled brownish-grey, nearly black on the hind-quarters, edged down the inside with reddish-yellow; the throat, belly, and legs lighter grey. Leith Adams ('P. Z. S.' 1858, p. 528) says: "Some are very dark on the upper parts, with black splashes on the back and hips; under-parts white or a dirty white. Others are of a yellowish-white all over the upper parts, with the belly and inner sides of the thighs white. A brownish-black variety is common, with a few white spots arranged longitudinally on the back—the latter I found were young." Kinloch writes: "The prevailing colour is brownish-grey, varying in shade on the back, where it is darkest, so as to give the animal a mottled or brindled appearance."
SIZE.—Length, about 3 feet; height, 22 inches.
The musk-deer is a forest-loving animal, keeping much to one locality. It bounds with amazing agility over the steepest ground, and is wonderfully sure-footed over the most rocky hills. It ruts in winter, produces one or two young, which are driven off in about six weeks' time by the mother to shift for themselves. They begin to produce at an early age—within a year. The musk bag is an abdominal or præputial gland which secretes about an ounce of musk, worth from ten to fifteen rupees. It is most full in the rutting season; in the summer, according to Leith Adams, it hardly contains any. The musk does not seem to affect the flavour of the meat, which is considered excellent.
CERVIDÆ—THE DEER.
CERVIDÆ—THE DEER.
Of the horned ruminants these are the most interesting. In all parts of the world, Old and New, save the great continental island of Australia, one or other kind of stag is familiar to the people, and is the object of the chase. The oldest writings contain allusions to it, and it is frequently mentioned in the Scriptures.
"Like as the hart desireth the water brooks,"
sang David. It is bound up in history and romance, and the chase of it in England is to this day a royal pastime.
However, to come back from the poetry of the thing to dry scientific details, I must premise that the two main distinctions of the Cervidæ, as separating them from the Bovidæ, are horns which are not persistent, but annually shed, and the absence of a gall bladder, which is present in nearly all the Bovidæ. The deer also, with one exception (the reindeer,Rangifer tarandus) have horns only in the males.
Regarding the shedding of these horns, it is supposed that the operation is connected with the sexual functions. It is a curious fact that castration has a powerful effect on this operation; if done early no horns appear; if later in life, the horns become persistent and are not shed.
Captain James Forsyth (in his 'Highlands of Central India'), was of opinion that the Sambar does not shed its horns annually, and states that this also is the opinion of native shikaris in Central India. This, however, requires further investigation. I certainly never heard of such a theory amongst them, nor noticed the departure from the normal state.
There have been several classifications of the Cervidæ, but I think the most complete and desirable one is that of Sir Victor Brooke (see'P. Z. S.' 1878, p. 883), which I shall endeavour to give in a condensed form. Dr. Gray's classification was based on three forms of antlers and the shape of the tail. But Sir Victor Brooke's is founded on more reliable osteological details. As I before stated in my introductory remarks on the Ruminantia, the first and fourth digits, there being no thumb, are but rudimentary, the metacarpal bones being reduced to mere splints; the digital phalanges are always in the same place, and bear the little false hoofs, which are situated behind and a little above the large centre ones, but the metacarpal splint is not always in the same place; it may either be annexed to the phalanges, or widely separated from them and placed directly under the carpus. The position of these splints is an important factor in the classification of the Cervidæ into two divisions, distinguished by Sir Victor Brooke as thePlesiometacarpals, in which the splint is near the carpus, and theTelemetacarpals, in which the splint is far from the carpus, and articulated with the digital phalanges. All the known species of deer can be classified under these two heads; and it is a significant fact that this pedal division is borne out by certain cranial peculiarities discovered by Professor Garrod, and also, to a certain extent, by an arrangement of hair-tufts on the tarsus and metatarsus. In the Old World deer, which are with few exceptionsPlesiometacarpi, those which have these tufts have them above the middle of the metatarsus, and those of the New World, which are, with one exception,Telemetacarpi, have them, when present, below the middle of the metatarsus.
There is also another character in addition to the cranial one before alluded to, which was also noticed by Professor Garrod. The first cranial peculiarity is that inTelemetacarpi, as a rule, the vertical plate developed from the lower surface of the vomer is prolonged sufficiently downwards and backwards to become anchylosed to the horizontal plate of the palatals, forming a septum completely dividing the nasal cavity into two chambers. In thePlesiometacarpithis vertical plate is not sufficiently developed to reach the horizontal plate of the palatals. The second cranial peculiarity is that in the Old World deer (Plesiometacarpi), the ascending rami of the premaxillæ articulate with the nasals with one or two exceptions, whereas in the New World deer (Telemetacarpi), with one or two exceptions, the rami of the premaxillæ do not reach the nasals. It will thus be seen that the osteological characters of the head and feet agree in a singularly fortunate manner, and, when taken in connection with the external signs afforded by the metatarsal tufts, prove conclusively the value of the system. In India we have to deal exclusively with thePlesiometacarpi, our nearest members of the other division being the Chinese water-deer (Hydropotes inermis), and probablyCapreolus pygargusfrom Yarkand, the horns of a roebuck in velvet attached to a strip of skin having been brought down by the Mission to that country in 1873-74.
Now comes the more difficult task of subdividing these sections into genera—a subject which has taxed the powers of many naturalists, and which is still in a far from perfect state. To all proposed arrangements some exception can be taken, and the following system is not free from objection, but it is on the whole the most reliable; and this system is founded on the form of the antler, which runs from a single spike, as in the South AmericanCoassus, to the many branches of the red deer (Cervus elaphas); and all the various changes on which we found genera are in successive stages produced in the red deer, which we may accept as the highest development; for instance, the stag in its first year develops but a single straight "beam" antler, when it is called a "brocket," and it is the same as the South American brocket (Coassus). On this being shed the next spring produces a small branch from the base of this beam, called the brow antler, which is identical almost with the single bifurcated horn of theFurciferfrom Chili. The stag is then technically known as a "spayad." In the third year an extra front branch is formed, known as the tres-tine. The antler then resembles the rusine type, of which our sambar stag is an example. In the fourth year the top of the main beam throws out several small tines called "sur-royals," and the brow antler receives an addition higher up called the "bez-tine." The animal is then a "staggard." In the fifth year the "sur-royals" become more numerous, and the whole antler heavier in the "stag," whose next promotion is to that of "great hart" of ten or more points. The finest heads are found in the German forests. Sir Victor Brooke alludes to some in the hunting Schloss of Moritzburg of the 15th to 17th century, of enormous size, bearing from 25 to 50 points—50 inches round the outside curve, 10 inches in circumference round thesmallestpart of the beam, and of one of which the spread between the coronal tines is 74 inches. Professor Garrod mentions one as having sixty-six points, and states that Lord Powerscourt has in his possession a pair with forty-five tines. The deer with which we have to deal range from the elaphine, or red deer type, to the simple bifurcated antler of the muntjac, which consists of a beam and brow antler only. We then come to the rusine type of three points only—brow, tres, and royal tines, and of this number are also the spotted and hog deer of India, but the arrangement of the tines is different; and following the rusine type comes the rucervine, in which the tres and royal tines break out into points—the tres-tine usually bifurcate, and the royal with two, three or more points. The arrangements of the main limbs of the horns is strictly rusine—that is to say, the external and anterior tine is equal to or shorter than the royal tine, whereas it is the reverse in the axis (spotted deer), and therefore this genus should come between the two. Even in the sambar and axis there is a tendency to throw out abnormal tines. There are many examples in the Indian Museum, and I possess a magnificent head which bears a large abnormal tine on one horn, and a faint inclination in the corresponding spot on the other horn to do likewise. I have no doubt, had the animal lived another year, the second extra tine would have been developed. Professor Garrod has three phases of the rucervine type, which he calls the normal, the intermediate, and the extreme. The first has both branches of the beam, tres and royal of equal size (ex. Schomburgk's deer); the second has the tres-tine larger than the royal (ex. our swamp deer); and the extreme type is that in which the royal is represented merely by a snag, the whole horn being bent forward (ex. the BurmesePanolia Eldii). The true cervine type of horn I have already described in its progress from youth to age. The Kashmir and Sikim stags are the representatives of this form in India. In Japan there is an intermediate form inCervus sikawhich has no bez-tine.