HUMIDITY

Can No.DateFormationStationCommunitySoilSampleHOLARDECHARDChresardNOTESWeighings%Weighings%SkyRainfall1st2dCan1st2dCan102/8/04Spruce forestJack BrookMertensiareLoam1058.750.125.52258.753.4125.521016Cloudy017„Spruce forestMilky WayGentianare¼ mold¾ gravel10:264.2557.521.351664.2559.621.35511Cloudy040„Gravel slide.HiawathaAsterareGravel2:1078.5574.322.85878.5574.8522.8517Cloudy0

A general designation of the soil composition is a material aid, especially where there is a difference in the core. For example, in a mountain forest or meadow, the upper layer will usually be mold, the lower sand or gravel. A careful estimate of the relation between the two throws much light upon the chresard. Under “sample” the number taken to reach the desired depth, if more than one, is indicated by placing the number before the depth, thus 2:10. When two or more full cores are included in the same sample for acheck, the order is 10:2. It has already been shown, however, that these precautions are not necessary for ordinary purposes. In computing the holard and echard, there is no need to show the figuring, if the process is checked and then proved. Notes upon sky conditions aid in explaining the daily decrease in water-content. The amount of rain and the period during which it falls are of great importance in understanding the fluctuations of the holard. Under community it is highly desirable to have a list of all the species, but it is impossible to include this in the table, and a glance at the formation list will show them. The form indicated above serves for a day-station series, a daily series in one station for any number of check series in one spot, and for scattered readings. In many cases the echard will not be determined, but on account of its primary importance, there should be a space for it, especially since it may be desirable to determine it at some later time.

56. The permanent record.This should be kept by formation, or if the latter exhibits well-defined associations, the formational record may be divided accordingly. This may seem an unnecessary expenditure of time, but a slight experience in finding the water-content values of a particular habitat, when scattered through a chronological field record, will be convincing. The form of permanent record is the same as for the field, except that the column for the formation and that for the society are often unnecessary.

57. Sums and means.From the great difficulty of determining the absolute water-content, and of obtaining a standard of comparison between soils on account of the varying ratio between bulk and weight, water-content sums are impracticable. For the same reasons, means of actual water-content are practically impossible, and the mean water-content must be expressed in per cents. Daily readings are not essential to a satisfactory mean. In fact, a single reading at each extreme enables one to approximate the real mean very closely; thus, the average of 26 readings in the prairie formation is 18 per cent. The extremes are 5 per cent and 28 per cent, and their average 16.5 per cent. A few readings properly scattered through moist and dry periods will give a reliable mean, as will also a series of daily readings from one heavy rain through a long dry period. The one difficulty with the last method is that such periods can not well be determined beforehand. Means permit ready comparison between habitats, but in connecting the modifications of a species with water-content as a cause, the extremes are significant as indicating the range of conditions. Furthermore, the extremes, i. e., 5 per cent and 28 per cent, make it possible to approximatethe mean, 18 per cent, while the latter gives little or no clue to the extremes. It is hardly necessary to state that means and extremes should be determined for a certain habitat, or particular area of it, and that the results may be expressed with reference to holard and chresard.

58. Curves.The value of graphic methods and the details of plotting curves are reserved for a particular section. It will suffice in this place to indicate the water-content curves that are of especial value. Simple curves are made with regard to time, place, or depth. The day curve shows the fluctuations of the water-content of one station from day to day or from time to time. The station curve indicates the variation in water from station to station, while the depth curve represents the different values at various depths in the same station. These may be combined on the same sheet in such a way that the station curves of each day may be compared directly. Similar combinations may be used for comparing the day curves, or the depth curves of different stations, but these are of less importance. A combination of curves which is of the greatest value is one which admits of direct comparison between the station curves of saturation, holard, chresard, and echard.

59. Instruments.As a direct factor, humidity is intimately connected with water-content in determining the structure and distribution of plants. The one is in control of water loss; the other regulates water supply. Humidity as a climatic factor undergoes greater fluctuation in the same habitat, and the efficient difference is correspondingly greater. Accordingly, simple instruments are less valuable than automatic ones, since a continuous record is essential to a proper understanding of the real influence of humidity. As is the rule, however, the use of simple instruments, when they can be referred to an ecographic basis, greatly extends the field which can be studied. In investigation, both psychrometer and psychrograph have their proper place. In the consideration of simple instruments for obtaining humidity values, an arbitrary distinction is made between psychrometers and hygrometers. The former consist of a wet and a dry bulb thermometer, while the latter make use of a hygroscopic awn, hair, or other object.

60. Kinds.There are three kinds of psychrometer, the sling, the cog, and the stationary. All consist of a wet bulb and a dry bulb thermometer set in a case; the first two are designed to be moved or whirled in the air. The same principle is applied in each, viz., that evaporation produces adecrease in temperature proportional to the amount of moisture in the air. The dry bulb thermometer is an ordinary thermometer, while the wet bulb is covered with a cloth that can be moistened. The former indicates the normal temperature of the air, the latter gives the reduced temperature due to evaporation. The relative humidity of the air is ascertained by means of the proper tables, from two terms, i. e., the air temperature and the amount of reduction shown by the wet bulb. The sling and the cog psychrometers alone are in general use. The stationary form has been found to be unreliable, because the moisture, as it evaporates from the wet bulb, is not removed, and, in consequence, hinders evaporation to the proper degree.

Fig. 5. Sling psychrometer.

61. The sling psychrometer.The standard form of this is shown in the illustration, and is the one used by the Weather Bureau. This instrument can be obtained from H. J. Green, 1191 Bedford Ave., Brooklyn, or Julien P. Friez, 107 E. German St., Baltimore, at a cost of $5. It consists of a metal frame to which are firmly attached two accurately standardized thermometers, reading usually from –30° to 130°. The frame is attached at the uppermost end to a handle in such fashion that it swings freely. The wet bulb thermometer is placed lower, chiefly to aid in wetting the cloth more readily. The cloth for the wet bulb should be always of the same texture and quality; the standard used by the Weather Bureau can be obtained from the instrument makers. A slight difference in texture makes no appreciable error, but the results obtained with different instruments and by different observers will be more trustworthy and comparable if the same cloth be used in all cases. The jacket for the wet bulb may be sewed in the form of a close-fitting bag, which soon shrinks and clings tightly. It may be made in the field by wrapping the cloth so that the edges just overlap, and tying it tightly above and below the bulb. In either case, a single layer of cloth alone must be used. The cloth becomes soiled or thin after a few months’ constant use and should be replaced. It is a wise precaution to carry a small piece of psychrometer cloth in the field outfit.

Fig. 6. Cog psychrometer.

62. Readings.All observations should be made facing the wind, and the observer should move one or two steps during the reading to prevent the possibility of error. The cloth of the wet bulb is moistened with water by means of a brush, or, much better, it is dipped directly into a bottle of water. Distilled water is preferable, as it contains no dissolved material to accumulate in the cloth. Tap-water and the water of streams may be used without appreciable error, if the cloth is changed somewhat more frequently. The temperature of the water is practically negligible under ordinary conditions. Readings can be made more quickly, however, when the temperature is not too far from that of the air. The psychrometer is held firmly and swung rapidly through the air when the space is not too confined. Where there is danger of breakage, it is swung back and forth through a short arc, pendulum-fashion. As the reading must be made when the mercury of the wet bulb reaches the lowest point, the instrument is stopped from time to time and the position of the column noted. The lowest point is often indicated by the tendency of the mercury to remain stationary; as a rule it can be noted with certainty when the next glance shows a rise in the column. In following the movement, and especially in noting the final reading, great care must be taken to make the latter before the mercury begins to rise. For this reason it is desirable to shade the psychrometer with the body when looking at it, and to take pains not to breathe upon the bulbs nor to bring them too near the body. At the moment when the wet bulb registers the lowest point, the dry bulb should be read and the results recorded.

63. Cog psychrometer.This instrument, commonly called the “egg-beater” psychrometer, has been devised to obviate certain disadvantages ofthe sling psychrometer in field work, and has entirely supplanted the latter in the writer’s own studies. It is smaller, more compact, and the danger of breaking in carriage or in use is almost nil. It has the great advantage of making it possible to take readings in a layer of air less than two inches in thickness, and in any position. Fairly accurate results can even be obtained from transpiring leaves. The instrument can readily be made by a good mechanic, at a cost for materials of $1.75, which is less than half the price for the sling form. A single drawback exists in the use of short, Centigrade thermometers, inasmuch as tables of relative humidity are usually expressed in Fahrenheit. It is a simple matter, however, to convert Centigrade degrees into Fahrenheit, mentally, or the difficulty may be avoided by the conversion table shown on page47, or by constructing a Centigrade series of humidity tables. The fact that the wet and dry bulbs revolve in the same path has raised a doubt concerning the accuracy of the results obtained with this instrument. Repeated comparisons with the sling psychrometer have not only removed this doubt completely, but have also proved that the standardization of the thermometers has been efficient.

64. Construction and use.A convenient form of egg-beater is the Lyon (Albany, New York), in which the revolving plates can be readily removed, leaving the axis and the frame. The thermometers used are of the short Centigrade type. They are 4½ inches long and read from –5° to 50°. Eimer and Amend, 205 Third Ave., New York city, furnish them at 75 cents each. The thermometers are carefully standardized and compared, and then grouped in pairs that read together. Each pair is used to construct a particular psychrometer. Each thermometer is strongly wired to one side of the frame, pieces of felt being used to protect the tube and increase the contact. The frame is also bent at the base angles to permit free circulation of air about the thermometer bulbs. The bulb of one thermometer is covered with the proper cloth, and the psychrometer is finished. Since the frame revolves with the thermometers, it is necessary to pour the water on the wet bulb, or to employ a pipette or brush. The thermometer bulbs are placed in the layer to be studied, and the frame rotated at an even rate and with moderate rapidity. The observation is further made as in the case of the sling psychrometer. As the circle of rotation is less than three inches in diameter, and the layer less than an inch, in place of nearly three feet for the sling form, the instrument should not be moved at all for extremely localized readings, but it must be moved considerably, a foot or more, if it is desirable to obtain a more general reading.

65. Hygrometers.While there are instruments designed to indicate the humidity by means of a hygroscopic substance, not one of them seems tobe of sufficient accuracy for use in ecological study. The difficulty is that the hygroscopic reaction is inconstant, rather than that the instruments are not sufficiently sensitive. A number of hygrometers have been tested, and in all the error has been found to be great, varying usually from 10–20 per cent. In the middle of the scale they sometimes read more accurately, but toward either extreme they are very inexact. It seems probable that an accurate hygrometer can be constructed only after the model of the Draper psychrograph. Its weight and bulk would make it an impossible instrument for field trips, and the expense of one would provide a dozen psychrometers. In consequence, it does not seem too sweeping to say that no hygrometer can furnish trustworthy results. Of simple instruments for humidity, the psychrometer alone can be trusted to give reliable readings. Crova’s hygrometer, used by Hesselmann, is not a hygrometer in the sense indicated. As it is much less convenient to handle and to operate than the cog psychrometer, it is not necessary to describe it.

Fig. 7. Draper psychrograph.

66. The Draper psychrograph.A year’s trial of the Draper psychrograph in field and planthouse has left little question of its accuracy and its great usefulness. Essentially, it consists of a band of fine catgut strings, which are sensitive to changes in the moisture-content of the air. The variations in the length of the band are communicated to a long pointer carrying an inking pen. The latter traces the record in per cent of relative humidity on a graduated paper disk, which is practically the face of an eight-day clock. The whole is enclosed in a metal case with a glass front. A glance at the illustration will show the generalstructure of the instrument. Continued psychrometric tests demonstrate that the margin of error is well within the efficient difference for humidity, which is taken to be 5 per cent. In the field tests of the past summer, two psychrographs placed side by side in the same habitat did not vary 1 per cent from each other. The same instruments when in different habitats did not deviate more than 1 per cent from the psychrometric values, except when the air approached saturation. For humidities above 90 per cent, the deviation is considerable, but as these are temporary and incident upon rainfall, the error is not serious. For humidities varying from 10–85 per cent, the psychrograph is practically as accurate as the psychrometer. Per cents below 10 are rare, and no tests have been made for them.

Fig. 8. Instrument shelter, showing thermograph and psychrograph in position.

67. Placing the instrument.The psychrograph should be located in a place where the circulation of the air is typical of the station observed. A satisfactory shelter will screen the instrument from sun and rain, and at the same time permit the air to pass freely through the perforations of the metal case. The form shown in figure 8 meets both of these conditions. A desirable modification is effected by fastening a strip about the cover of such depth as to prevent the sun’s rays from striking the case except when the sun is near the horizon. A cross block is fastened on thepost of the shelter after being exactly leveled. The psychrograph rests upon this block, which is three feet above the ground in order to avoid the influence of radiation. The instrument is held in position by slipping the eye over a small-headed nail driven obliquely. It does not hang from the latter, but must rest firmly upon the cross block. The post is set to a depth that prevents oscillation in the wind, which is liable to obscure the record. In shallow mountain soils stability is attained by fastening a broad board at the base of the post before setting it. When two or more psychrographs are established in different habitats, great pains are taken to set them up in exactly the same way. The shelters are alike, the height above the soil the same, and the instruments all face the south.

68. Regulating and operating the instrument.When two or more psychrographs are to be used in series, they must be compared with each other in the same spot for several days until they run exactly together with respect to per cent of humidity and to time. During this comparison they are checked by the psychrometer and so regulated that they register the proper humidity. When a single instrument is used alone as the basis to which simple readings may be referred, all regulating may well be done after the instrument is in position. This is a simple process; it is accomplished by obtaining the relative humidity beneath the shelter and at the proper height by a psychrometer. The pen hand is then moved to the proper line on the disk by means of the screws at its base. These are reached by removing the lettered glass face. The thumbscrew on the side opposite the direction in which the pen is to move is released, and the opposite screw simultaneously tightened, until the pen remains upon the proper line. Experience has proved that the record sheet should be correctly labeled and dated before being placed on the disk. In the press of field duties, records labeled after removal are liable to be confused. It is likewise a great saving of time to write the date of the month in the margin of each segment. Care is taken to place the sheet on the disk in the same position each time; this can easily be done by seeing that the sharp point on the disk penetrates the same spot on the paper. A single drop of ink in the pen will usually give the most satisfactory line. A thin line is read most accurately. If the pen point is too fine, however, the ink does not flow readily, and the point should be slightly blunted by means of a file. More often the line is too broad and the pen must be carefully pointed. Occasionally the pen does not touch the sheet, and it becomes necessary to bend the hand slightly. This is a frequent difficulty if the records are folded or wrinkled, and consequently the sheets should always be kept flat.

69. The weekly visit.Psychrographs must be visited, checked, rewound, and inked every week. Whenever possible this should be done regularly at a specified day and hour. This is especially desirable if the same record sheet is used for more than one week. Time and energy are saved by a fixed order for the various tasks to be done at each visit. After opening the instrument the disk is removed, and the clock wound, and, if need be, regulated. The record sheet is replaced, the disk again put on the clock arbor, and the pen replenished with a drop of ink. A psychrometer reading is made, and the results in terms of relative humidity noted at the proper place on the disk sheet. If the psychrograph vary more than 1 per cent, it is adjusted to read accurately. In practice it has been found a great convenience to keep each record sheet in position for three weeks, and the time may easily be extended to four. In this event, the pen is carefully cleaned with blotting paper at each visit, and is then refilled with an ink of different color. To prevent confusion, the three different colored inks are always used in the same order, red for the first week, blue for the second, and green for the third. The advantages of this plan are obvious: fewer records are used and less time is spent in changing them. The records of several weeks are side by side instead of on separate sheets, and in working over the season’s results, it is necessary to handle but a third as many sheets.

The Draper psychrograph is made by the Draper Manufacturing Company, 152 Front St., New York city. The price is $30. A few record sheets and a bottle of red ink are furnished with it. Additional records can be obtained at 3 cents each. The inks are 25–50 cents per bottle, depending upon the color.

70. The time of readings.If simple instruments alone are used for determining humidity, readings are practically without value unless made simultaneously through several stations, or successively at one. When it is possible to combine these, and to make psychrometer readings at different habitats for each hour of the day, or at the same hour for several days, the series is of very great value. Single readings are unreliable on account of the hourly and daily variations of humidity, but when these changes are recorded by a psychrograph, such readings at once become of use, whether made in the same habitat with the recording instrument or elsewhere. In the latter case, one reading will tell little about the normal humidity of the habitat, but several make a close estimate possible. When a series of psychrographs is in use, accurate observations can be made to advantage anywhere at any time. As a rule, however, it has been found most convenientto make simple readings at 6:00A.M., 1:00P.M., and 6:00P.M., as these hours afford much evidence in regard to the daily range. A good time also is that at which the temperature maximum occurs each day, but this is movable and in the press of field work can rarely be taken advantage of. A very fair idea of the daily mean humidity is obtainable by averaging the readings made at the hours already indicated. The comparison of single readings with the psychrograph record should not be made at a time when a rapid change is occurring, as the automatic instrument does not respond immediately. Such a condition is usually represented by a sudden rain, and is naturally not a satisfactory time for single readings in any event.

Fig. 9. Atmometer.

71. Place and height.As stated above, the psychrograph is placed three feet above the surface of the ground in making readings for the comparison of stations. In low, herbaceous formations, the instrument is usually placed within a few inches of the soil in order to record the humidity of the air in which the plants are growing. In forest formations, the moisture often varies considerably in the different layers. This variation is easily determined by simultaneous psychrometer readings in the several layers, or, if occasion warrants, a series of psychrographs may be used. In field work the rule has been to make observations with the psychrometer at 6 feet, 3 feet, and the surface of the soil, but the reading at the height of 3 feet is ordinarily sufficient. Humidity varies so easily that several readings in different parts of one formation are often desirable. In comparing different formations, the readings should be made in corresponding situations, for example, in the densest portion of each.

72. Check instruments.Humidity is so readily affected by temperature, wind, and pressure, that a knowledge of these factors is essential to an understanding of its fluctuations. Pressure, disregarding daily variation, is taken account of in the tables for ascertaining relative humidity, and is determined once for all when the altitude of a station has been carefully established. The temperature is obtained directly from the dry bulb reading. Its value is fundamental, as the amount of moisture in a given space is directly affected by it; like pressure, it also is taken account of in the formula. The movement of the air has an immediate influenceupon moisture by mixing the air of different habitats and layers. So far as the plant is concerned, it has practically the effect of increasing or decreasing the humidity by the removal of the air above it. Thus, while the anemometer can furnish no direct evidence as to the amount of variation, it is of aid in explaining the reason for it. Likewise, the rate of evaporation as indicated by a series of atmometers, affords a ready method of estimating the comparative effect of humidity in different habitats. Potometers and other instruments for measuring transpiration throw much light upon humidity values. Since they are concerned with the response of the plant to humidity, they are considered in the following chapter.

73. Humidity tables.To ascertain the relative humidity, the difference between the wet and dry bulb readings is obtained. This, with the dry bulb temperature, is referred to the tables, where the corresponding humidity is found. A variation in temperature has less effect than a variation in the difference; in consequence, the dry bulb reading is expressed in the nearest unit, and the difference reckoned to the nearest .5. The humidity varies with the air pressure. Hence, the altitude must be determined for the base station, and for all others that show much change in elevation. Within the ordinary range of growing-period temperatures, the effect of pressure is not great. For all ordinary cases, it suffices to compute tables for pressures of 30, 29, 27, 25, and 23 inches. The following table indicates the decrease in pressure which is due to altitude.

The fluctuations of pressure due to weather are usually so slight that their influence may be disregarded. An excellent series of tables of relative humidity is found in Marvin’s Psychrometric Tables, published by the U. S. Weather Bureau, and to be obtained from the Division of Publications, Washington, D. C., for 10 cents. A convenient field form is made by removing the portion containing the tables of relative humidity, and binding it in stiff oilcloth.

Fig. 10. Conversion scale for temperatures.

74. Sums, means, and curves.An approximate humidity sum can be obtained by adding the absolute humidities for each of the twenty-four hours, and expressing the results in grains per cubic foot. It is possible to establish a general ratio between this sum and the transpiration sum of the plant, but its value is not great at present. Means of absolute and of relative humidity are readily determinable from the psychrograph records; the latter are the most useful. The mean of relative humidity for the twenty-four hours of a day is the average of the twenty-four hour humidities. From these means the seasonal mean is computed in the same manner. A close approximation, usually within 1 degree, may be obtained in either case by averaging the maximum and minimum for the period concerned. Various kinds of curves are of value in representing variation in humidity. Obviously, these must be derived from the psychrograph, or from the psychrometer when the series is sufficiently complete. The level curve indicates the variation in different stations at the same time.These may be combined in a series for the comparison of readings made at various heights in the stations. The day or point curve shows the fluctuations during the day of one point, and the station curve the variation at different heights in the same station. The curves of successive days or of different stations may of course be combined on the same sheet for comparison. Level and station curves based upon mean relative humidities are especially valuable.

75. Records.A field form is obviously unnecessary for the psychrograph. The record sheets constitute both a field and permanent record. The altitude and other constant features of the station and the list of species, etc., are entered on the back of the first record sheet, or, better, they are noted in the permanent formation record. For psychrometer readings, whether single or in series, the following record form is employed:

DateHourFormationStationAltitudeCommunityHeight of readingDry bulbWet bulbDiff.Rel. Hum.Base Hum.Abs. Hum.NOTESSkyRainWind15/8/’046:20A.M.SpruceBrook bank2500 mMertensiare1 ft.51°46°572%63%2.9Clear00„„Half gravelHiawatha„Asterare„56°49°764%63%3.0„00„6:45P.M.SpruceBrook bank„Mertensiare„54°52°289%69%4.2„2 cc.0„„Half gravelHiawatha„Asterare„56°52°479%69%4.0„2 cc.0

On page47is given a table for the conversion of Centigrade into Fahrenheit temperatures. This may be done mentally by means of the formulaF=C/5 × 9 + 32°.

76. Methods.All methods for measuring light intensity, which have been at all satisfactory, are based upon the fact that silver salts blacken in the light. The first photographic method was proposed by Bunsen and Roscoe in 1862; this has been taken up by Wiesner and variously modified. After considerable experiment by the writer, however, it seemed desirable to abandon all methods which require the use of “normal paper” and “normal black” and to develop a simpler one. As space is lacking for a satisfactory discussion of the Bunsen-Roscoe-Wiesner methods, the reader is referred to the works cited below.[4]Simple photometers for making light readings simultaneouslyor in series were constructed in 1900, and have been in constant use since that time. An automatic instrument capable of making accurate continuous records proved to be a more difficult problem. A sunshine recorder was ultimately found which yields valuable results, and very recently a recording photometer which promises to be perfectly satisfactory has been devised. Since the hourly and daily variations of sunlight in the same habitat are relatively small, automatic photometers are perhaps a convenience rather than a necessity.

Fig. 11. Photometer, showing front and side view.

77. Construction.The simple form of photometer shown in the illustration is a light-tight metal box with a central wheel upon which a strip of photographic paper is fastened. This wheel is revolved by the thumbscrew past an opening 6 mm. square which is closed by means of a slide working closely between two flanges. At the edge of the opening, and beneath the slide is a hollow for the reception of a permanent light standard. The disk of the thumbscrew is graduated into twenty-five parts, and these are numbered. A line just beneath the opening coincides withthe successive lines on the disk, and indicates the number of the exposure. The wheel contains twenty-five hollows in which the click works, thus moving each exposure just beyond the opening. The metal case is made in two parts, so that the bottom may be readily removed, and the photographic strip placed in position. The water-photometer is similar except that the opening is always covered with a transparent strip and the whole instrument is water-tight. These instruments have been made especially for measuring light by the C. H. Stoelting Co., 31 W. Randolph street, Chicago, Ill. The price is $5.

78. Filling the photometer.The photographic paper called “solio” which is made by the Eastman Kodak Company, Rochester, N. Y., has proved to be much the best for photometric readings. The most convenient size is that of the 8 × 10 inch sheet, which can be obtained at any supply house in packages of a dozen sheets for 60 cents. New “emulsions,” i. e., new lots of paper, are received by the dealers every week, but each emulsion can be preserved for three to six months without harm if kept in a cool, light-tight place. Furthermore, all emulsions are made in exactly the same way, and it has been impossible to detect any difference in them. To fill the photometer, a strip exactly 6 mm. wide is cut lengthwise from the 8 × 10 sheet. This must be done in the dark room, or at night in very weak light. The strip is placed on the wheel, extreme care being taken not to touch the coated surface, and fixed in position by forcing the free ends into the slit of the wheel by a piece of cork 8–9 mm. long. The wheel is replaced in the case, turned until the zero is opposite the index line, and the instrument is ready for use.

79. Making readings.An exposure is made by moving the slide quickly in such a way as to uncover the entire opening, and the standard if the exposure is to be very short. Care must be taken not to pull the slide entirely out of the groove, as it will be impossible to replace it with sufficient quickness. The time of exposure can be determined by any watch after a little practice. It is somewhat awkward for one person to manage the slide properly when his attention is fixed upon a second hand. This is obviated by having one observer handle the watch and another the photometer, but here the reaction time is a source of considerable error. The most satisfactory method is to use a stop-watch. This can be held in the left hand and started and stopped by the index finger. The photometer is held against it in the right hand in such a way that the two movements of stopping the watch and closing the slide may be made at the same instant. The length of exposure is that necessary to bring the tint of the paper to that of thestandard beside it. A second method which is equally advantageous and sometimes preferable does away with the permanent standard in the field and the need for a stop-watch. In this event, the strip is exposed until a medium color is obtained, since very light or very deep prints are harder to match. This is later compared with the multiple standard. In both cases, the date, time of day, station, number of instrument and of exposure, and the length of the latter in seconds are carefully noted. The instrument is held with the edge toward the south at the level to be read, and the opening uppermost in the usual position of the leaf. When special readings are desired, as for isophotic leaves, reflected light, etc., the position is naturally changed to correspond. In practice, it is made an invariable rule to move the strip for the next exposure as soon as the slide is closed. Otherwise double exposures are liable to occur. When a strip is completely exposed it is removed in the dark, and a new one put in place. The former is carefully labeled and dated on the back, and put away in a light-tight box in a cool place.

Fig. 12. Dawson-Lander sun recorder.

80. The Dawson-Lander sun recorder.“The instrument consists of a small outer cylinder of copper which revolves with the sun, and through the side of which is cut a narrow slit to allow the sunshine to impinge on a strip of sensitive paper, wound round a drum which fits closely inside the outer cylinder, but is held by a pin so that it can not rotate. By means ofa screw fixed to the lid of the outer cylinder, the drum holding the sensitive paper is made to travel endwise down the outer tube, one-eighth of an inch daily, so that a fresh portion of the sensitive surface is brought into position to receive the record.” The instrument is driven by an eight-day clock placed in the base below the drum. The slit is covered by means of a flattened funnel-shaped hood, and the photographic strip is protected from rain by a perfectly transparent sheet of celluloid. The detailed structure of the instrument is shown in figure 12. This instrument may be obtained from Lander and Smith, Canterbury, England, for $35.

In setting up the sunshine recorder, the axis should be placed in such a position that the angle which it makes with the base is the same as the altitude of the place where the observations are made. This is readily done by loosening the bolts at either side. The drum is removed, the celluloid sheet unwound by means of the key which holds it in place, the sensitive strip put in position, and the sheet again wound up. Strips of a special sensitive paper upon which the hours are indicated are furnished by the makers of the instrument, but it has been found preferable to use solio strips in order to facilitate comparison with the standards. The drum is placed on the axis, and is screwed up until it just escapes the collar at the top of the spiral. The clock is wound and started, and the outer cylinder put on so that the proper hour mark coincides with the index on the front of the base.

As a sunshine recorder, the instrument gives a perfect record, in which the varying intensities are readily recognizable. Since the cylinder moves one-half inch in an hour, and the slit is .01 of an inch, the time of each exposure is 72 seconds. This gives a very deep color on the solio paper, which results in a serious error in making comparisons with the standard. On account of the hood, diffuse light is not recorded when it is too weak to cast a distinct shadow. It seems probable that this difficulty will be overcome by the use of a flat disk containing the proper slit, and in this event the instrument will become of especial value for measuring the diffuse light of layered formations. The celluloid sheet constitutes a source of error in sunlight on account of the reflection which it causes. This can be prevented by using the instrument only on sunny days, when the protection of the sheet can be dispensed with.

81. The selagraph.This instrument is at present under construction, and can only be described in a general way. In principle it is a simple photometer operating automatically. It consists of a light-tight box preferably of metal, which contains an eight-day lever clock. Attached to the arbor of the latter is a disk 7 inches in diameter bearing on its circumference a solio strip 1 cm. wide and 59 cm. long. The opening in the boxfor exposure is 6 mm. square and is controlled by a photographic shutter. The latter is constructed so that it may be set for 5, 10, or 20 seconds, since a single period of exposure can not serve for both sun and shade. The shutter is tripped once every two hours, by means of a special wheel revolving once a day. Each exposure is 6 mm. square, and is separated by a small space from the next one. Twelve exposures are made every 24 hours, and 84 during the week, though, naturally, the daytime exposures alone are recorded. Comparisons with the multiple standard are made exactly as in the case of the simple photometer. The selagraph is made by the C. H. Stoelting Co., Chicago, Illinois.

82. Use.The light value of each exposure is determined by reference to a standard. When the photometer carries a permanent standard, each exposure is brought to the tint of the latter, and its value is indicated by the time ratio between them. Thus, if the standard is the result of a 5–second exposure to full sunlight at meridian, and a reading which corresponds in color requires 100 seconds in the habitat concerned, the light of the latter is twenty times weaker or more diffuse. Usually, the standard is regarded as unity, and light values figured with reference to it, as .05. With the selagraph such a use of the standard is impossible, and often, also, with the photometer it is unnecessary or not desirable. The value of each exposure in such case is obtained by matching it with a multiple standard, after the entire strip has been exposed. The further steps are those already indicated. After the exact tint in the standard has been found, the length of the reading in seconds is divided by the time of the proper standard, and the result expressed as above.

83. Making a standard.Standards are obtained by exposing the photometer at meridian on a typically clear day, and in the field where there is the least dust and smoke. Exception to the latter may be made, of course, in obtaining standards for plant houses located in cities, though it is far better to have the same one for both field and control experiment. Usable standards can be obtained on any bright day at the base station. Indeed, valuable results are often secured by immediate successive sun and shade readings in adjacent habitats, where the sun reading series is the sole standard. Preferably, standards should be made at the solstices or equinoxes, and at a representative station. The June solstice is much to be preferred, as it represents the maximum light values of the year. Lincoln has been taken as the base station for the plains and mountains. It is desirable, however,that a national or international station be ultimately selected for this purpose, in order that light values taken in different parts of the world may be readily compared.

84. Kinds of standards.The base standard is the one taken at Lincoln (latitude 41° N.) at meridian June 20–22. This is properly the unit to which all exposures are referred, but it has been found convenient to employ the Minnehaha standard as the base for the Colorado mountains, in order to avoid reducing each time.Relativestandards are frequently used for temporary purposes. Thus, in comparing the light intensities of a series of formations, one to five standards are exposed on the solio strip before beginning the series of readings.Proofstandards are the exposed solio strips, which fade in the light, and can, in consequence, be kept only a few weeks without possibility of error. The fading can be prevented by “toning” the strip, but in this event the exposures must be fixed in like manner before they can be compared. This process is inconvenient and time-consuming. It is also open to considerable error, as the time of treatment, strength of solution, etc., must be exactly equivalent in all instances.Permanentstandards are accurate water-color copies of the originals obtained by the photometer. These have the apparent disadvantage of requiring a double comparison or matching, but after a little practice it is possible to reproduce the solio tints so that the copy is practically indistinguishable from the original. The most satisfactory method is to make a long stroke of color on a pure white paper, since a broad wash is not quite homogeneous, and then to reject such parts of the stroke as do not match exactly. Permanent standards fade after a few month’s use, and must be replaced by parts of the original stroke.Singlestandards are made by one exposure, whilemultipleones have a series of exposures filling a whole light strip. These are regularly obtained by making the exposures from 1–10 seconds respectively, and then increasing the length of each successive exposure by 2 seconds. Single exposures of 1–5 seconds as desired usually serve as the basis for permanent standards, but a multiple standard may also be copied in permanent form. Exposures for securing standards must be made only under the most favorable conditions, and the length in seconds must be exact. The use of the stop-watch is imperative, except where access may be had to an astronomical clock with a large second hand, which is even more satisfactory. The length of time necessary for the series desired is reckoned beforehand, and the exposures begun so that the meridian falls in the middle of the process.

Single standards are exceedingly convenient in photometer readings, but they are open to one objection. In the sunshine it is necessary to make instant decision upon the accuracy of the match, or the exposure becomes toodeep. In the shade where the action is slower, this difficulty is not felt. For this reason it is usually desirable to check the results by a multiple standard, and in the case of selagraph records, where the various exposures show a wide range of tint, light values are obtainable only by direct comparison with the multiple standard. The exact matching of exposure and standard requires great accuracy, but with a little practice this may be done with slight chance of error by merely moving the exposure along the various tints of the standard until the proper shade is found. The requisite skill is soon acquired by running over a strip of exposures several times until the comparisons always yield the same results for each. The margin of error is practically negligible when the same person makes all the comparisons, and in the case of two or three working on the same reading the results diverge little or not at all. The efficient difference for light is much more of a variable than is the case with water-content. It has been determined so far only for a few species, all of which seem to indicate that appreciable modification in the form or structure of a leaf does not occur until the reduction in intensity reaches .1 of the meridian sunlight at the June solstice. The error of comparison is far less than this, and consequently may be ignored, even in the most painstaking inquiry.

85. Time.The intensity of the light incident upon a habitat varies periodically with the hour and the day, and changes in accord with the changing conditions of the sky. The light variations on cloudy days can only be determined by the photometer. While these can not be ignored, proper comparisons can be instituted only between the readings taken on normal days of sunshine. The sunlight varies with the altitude of the sun, i. e., the angle which its rays make with the surface at a given latitude. This angle reaches a daily maximum at meridian. The yearly maximum falls on June 22, and the angle decreases in both directions through the year to a minimum on December 22. At equal distances from either solstice, the angle is the same, e. g., on March 21 and September 23. At Lincoln (41° N. latitude) the extremes at meridian are 73° and 26°; at Minnehaha (39°) they are 75° and 28°. The extremes for any latitude may be found by subtracting its distance in degrees north of the two tropics from 90. Thus, the 50th parallel is 26.5° north of the tropic of Cancer, and the maximum altitude of the sun at a place upon it is 63.5°. It is 73.5° north of the tropic of Capricorn, and the minimum meridional altitude is 16.5°.

The changes in the amount of light due to the altitude of the sun are produced by the earth’s atmosphere. The absorption of light rays is greatest near the horizon, where their pathway through the atmosphere is longest,and it is least at the zenith. The absorption, and, consequently, the relative intensity of sunlight, can be determined at a given place for each hour of any sunshiny day by the use of chart 13. This chart has been constructed for Lincoln, and will serve for all places within a few degrees of the 40th parallel. The curves which show the altitude of the sun at the various times of the day and the year have been constructed by measurements upon the celestial globe. Each interval between the horizontal lines represents 2 degrees of the sun’s altitude. The vertical lines indicate time before or after the apparent noon, the intervals corresponding to 10 minutes. If the relative intensity at Lincoln on March 12 at 3:00P.M.is desired, the apparent noon for this day must first be determined. A glance at the table shows that the sun crosses the meridian on this day at 9 minutes 53 seconds past noon at the 90th meridian. The apparent noon at Lincoln is found by adding 26 minutes 49 seconds, the difference in time between Lincoln and a point on the 90th meridian. When the sun is fast, the proper number of minutes is taken from 26 minutes 49 seconds. The apparent noon on March 12 is thus found to fall at 12:37P.M., and 3:00P.M.is 2 hours and 23 minutes later. The sun’s altitude is accordingly 36°. If the intensity of the light which reaches the earth’s surface when the sun is at zenith is taken as 1, the table of the sun’s altitudes gives the intensity at 3:00P.M.on March 12 as .85.

For places with a latitude differing by several degrees from that of Lincoln, it is necessary to construct a new table of altitude curves from the celestial globe. It is quite possible to make a close approximation of this from the table given, since the maximum and minimum meridional altitude, and hence the corresponding light intensity, can be obtained as indicated above. For Minnehaha, which is on the 105th meridian, and for other places on standard meridians, i. e., 60°, 75°, 90°, and 120° W., the table of apparent noon indicates the number of minutes to be added to 12 noon, standard time, when the sun is slow, and to be subtracted when the sun is fast. The time at a place east or west of a standard meridian is respectively faster or slower than the latter. The exact difference in minutes is obtained from the difference in longitude by the equation, 15° = 1 hour. Thus, Lincoln, 96° 42′ W. is 6° 42′ west of the standard meridian of 90°; it is consequently 26 minutes 49 seconds slower, and this time must always be added to the apparent noon as determined from the chart. At a place east of a standard meridian, the time difference is, of course, subtracted.

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