Dipodomys ordii nexilisGoldman, Journ. Washington, Acad. Sci., 23:470, October 5, 1933.
Dipodomys ordii nexilisGoldman, Journ. Washington, Acad. Sci., 23:470, October 5, 1933.
Perodipus longipes,Warren, Mammals of Colorado, p. 77, 1910 (part—the part from Coventry, Colorado).
Perodipus longipes,Warren, Mammals of Colorado, p. 77, 1910 (part—the part from Coventry, Colorado).
Dipodomys ordii longipes, Moore, Journ. Mamm., 10:260, August, 1929 (part—the part from Monticello, Utah).
Dipodomys ordii longipes, Moore, Journ. Mamm., 10:260, August, 1929 (part—the part from Monticello, Utah).
Type.—Male, adult, no. 149938, U. S. Nat. Mus. Biol. Surv. Coll.; 5 mi. W Naturita, Montrose County, Colorado; obtained on July 20, 1907, by Merritt Cary, original no. 1068.
Range.—West-central Colorado, southwest into San Juan County, Utah, north of the San Juan River; northwest into Grand County, Utah, to theColorado River; and westward probably as far as the Colorado River; marginal occurrences are: in Utah, Cisco, 18 mi. NE Moab, and Blanding; in Colorado, 5 mi. W Naturita, and Coventry.
Diagnosis.—Size medium (see measurements). Color dark, entire dorsal surface between (a) Cinnamon-Buff and Clay Color, purest on sides and flanks, upper parts strongly suffused with black; arietiform markings, pinnae of ears, plantar surfaces of hind feet, dorsal and ventral tail-stripes blackish. Lateral white stripes of tail less than one fourth of diameter of tail. Skull large; rostrum long and narrow; nasals long; auditory bullae strongly inflated; braincase not inflated; pterygoid fossae ovoid.
Comparisons.—FromDipodomys ordii sanrafaeli,D. o. nexilisdiffers in: Color darker; skull smaller in all measurements taken; auditory bullae noticeably smaller; width across zygomatic processes of maxillae less.
FromDipodomys ordii longipes,D. o. nexilisdiffers in: Size smaller; color darker in all respects; rostrum wider; breadth across auditory bullae less; interorbital width greater; breadth across zygomatic processes of maxillae less; cutting edge of upper incisors wider; zygomatic arch lighter and more nearly straight.
Comparison withDipodomys ordii evexusis made in account of that subspecies.
Remarks.—This animal is apparently not abundant at any place in its range. Two different attempts, by me, to obtain topotypes were unsuccessful. A single specimen was obtained and that was only a skull saved from a mutilated animal that was taken away from a rattlesnake. The habitat at the type locality is such that a person would not expect it to be inhabited by kangaroo rats. The soil is a heavy clay with a generous admixture of stones but in isolated spots there are light sandy soils which should be suitable for kangaroo rats. Even so, 500 traps set in the area of the type locality over a period of two nights yielded noDipodomys.
This subspecies ofDipodomys ordii, inhabiting west-central Colorado and southeastern Utah, is darker than any other subspecies with which its range comes in contact. The races to both the north and south are larger, with the exception of the hind foot which is longer inD. o. nexilisthan in eitherD. o. sanrafaeliorD. o. longipes. The affinities ofD. o. nexilisare withD. o. longipesrather than with any of the other known subspecies ofDipodomys ordii.
Intergradation betweenD. o. nexilisandD. o. longipesis shown by animals from Bluff, San Juan County, Utah, which, however, are referable to the latter. In animals from sixteen miles northwest of Moab, Grand County, Utah, there is intergradation in size of skull and in color betweenD. o. nexilisandD. o. sanrafaeli. The specimens are referred toD. o. sanrafaeli.
D. o. nexilisis apparently prevented from extending its range to the northward by the presence there ofD. o. sanrafaeli. To the southD. o. nexilisencountersD. o. longipesand the San Juan River. To the east it is limited by the Rocky Mountains and to the west by the deep canyons of the Colorado River.
Warren(1942:183) did not apply the nameD. o. nexilisto Ord kangaroo rats in Colorado. He listed specimens from Montrose, Montezuma and Mesa counties as being referable toD. o. longipes. The specimens from Montezuma County, Colorado, probably areD. o. longipesand those from Montrose and Mesa counties (for whichWarrengives no precise localities) areD. o. nexilis.
Specimens examined.—Total, 35, distributed as follows:
Utah:Grand County: Cisco, 4 (CM); 18 mi. NE Moab, 6000 ft., 1 (UU).San Juan County: 15 mi. N Monticello, 4 (MVZ); Blanding, 1 (UU).
Colorado:Delta County: Hotchkiss, 1 (USBS).Montrose County: Paradox, 1 (DRD); Bedrock, 4 (AMNH); 5 mi. W Naturita, 2 (USBS); Coventry, 3 (1 USBS; 2 AMNH).
Dipodomys ordii cupidineusGoldman, Journ. Washington Acad. Sci., 14:372, September 19, 1924.
Dipodomys ordii cupidineusGoldman, Journ. Washington Acad. Sci., 14:372, September 19, 1924.
Type.—Male, no. 243093, U. S. Nat. Mus. Biol. Surv. Coll.; Kanab Wash, at southern boundary of Kaibab Indian Reservation, Arizona; obtained on October 12, 1922, by E. A.Goldman, original no. 23384.
Range.—Northeastern Arizona and south-central Utah; marginal occurrences are: in Utah, Escalante, near Paria and S of Virgin; in Arizona, 6 mi. N Wolf Hole, 20 mi. S Wolf Hole, 5 mi. S Trumbull Spring and 10 mi. S Jacobs Pools in Houserock Valley.
Diagnosis.—Size large (see measurements). Color relatively dark, entire dorsal surface between (16") Pinkish Cinnamon and Cinnamon-Buff, purest on sides and flanks, upper parts washed with black; arietiform markings, pinnae of ears, dorsal and ventral stripes of tail, plantar surfaces of hind feet, blackish. Skull large; auditory bullae well inflated; interorbital width relatively narrow; rostrum long and narrow; nasals long and slightly flared distally; pterygoid fossae ovoid; external auditory meatus small and ovoid; jugal light and relatively straight.
Comparisons.—FromDipodomys ordii chapmani,D. o. cupidineusdiffers in: Size larger; color lighter in all pigmented areas; skull larger; rostrum shorter and wider; nasals shorter; auditory bullae more inflated, particularly median tympanic portion; least interorbital width less.
FromDipodomys ordii nexilis,D. o. cupidineusdiffers as follows: Size smaller; color lighter in all pigmented areas; skull smaller; rostrum relatively longer and narrower; least interorbital width less; auditory bullae less inflated; breadth across maxillary arches less; foramen magnum circular as opposed to ovoid; cutting edge of upper incisors narrower; zygomatic arches lighter.
For comparisons withDipodomys ordii cinderensis,Dipodomys ordii panguitchensisandDipodomys ordii longipessee accounts of those subspecies.
Remarks.-This subspecies is prevented from intergrading, to the east, withD. o. longipesby the deep canyons of the Colorado River, withD. o. cinderensisby the Virgin River and withD. o. panguitchensisby a series of high mountain ranges. Intergradation does, however, occur withD. o. sanrafaeliin animals from Escalante, Garfield County, Utah, but they are referable toD. o. cupidineus. The Colorado River, farther to the north, again serves as a barrier in preventing intergradation betweenD. o. nexilisandD. o. cupidineus.
Specimens examined.—Total, 122, distributed as follows:
Utah:Garfield County: Mouth of Calf Creek, Escalante River, 3 (BYU); Ten Mile Spring, 3 (BYU); Escalante, 3 (BYU).Washington County: near Short Creek road, south of town of Virgin, 18 (RH).Kane County: near Paria, 1 (UU); Willow Tank Springs, 7 (BYU); 8 mi. NW Kanab, 4800 ft., 1 (UU); near Sand Dunes, 7 (RH); Kanab, 15 (12 MVZ; 2 BYU; 1 UU); 1 mi. S Kanab, 4400 ft., 2 (UU).
Arizona:Mohave County: near S Boundary Kaibab Indian Reservation, 4 (MVZ); 6 mi. N Wolf Hole, 8 (MVZ); 4 mi. N Wolf Hole; 3 mi. NW Diamond Butte, 7 (USBS); 20 mi. S Wolf Hole, 1 (USBS); 1 mi. W Diamond Butte, 1 (USBS); 6 mi. N Mt. Trumbull, 1 (USBS); Nixon Springs, 6250 ft., W Base Mt. Trumbull, 6 (3 USBS; 3 MVZ); 5 mi. S Trumbull Spring, 1 (USBS).Coconino County: Fredonia, 3 (USBS); 2 mi. W Lees Ferry, 3 (MVZ); Jacobs Pools, 4000 ft., Houserock Valley, 15 (MVZ); 10 mi. S Jacobs Pools, Houserock Valley, 2 (MVZ); Houserock Valley, 10 (USBS).
Dipodops ordii palmeriAllen, Bull. Mus. Comp. Zool., 8:187, 1881.
Dipodops ordii palmeriAllen, Bull. Mus. Comp. Zool., 8:187, 1881.
Cricetodipus ordii palmeri, Trouessart, Catalogus Mammalium, 1:581, 1897.
Cricetodipus ordii palmeri, Trouessart, Catalogus Mammalium, 1:581, 1897.
Perodipus ordii palmeri,Goldman, Proc. Biol. Soc. Washington, 30:113, May 23, 1917.
Perodipus ordii palmeri,Goldman, Proc. Biol. Soc. Washington, 30:113, May 23, 1917.
Dipodomys ordii palmeri,Grinnell, Journ. Mamm., 2:96, May 2, 1921.
Dipodomys ordii palmeri,Grinnell, Journ. Mamm., 2:96, May 2, 1921.
Cotypes.—Two males, adults, nos. 5886 and 5887, Mus. Comp. Zool.; San Luis Potosí, Mexico; obtained on May 1, 1878, and September 1, 1878, respectively, by Dr. Edward Palmer. (Types not seen.)
Range.—Eastern Zacatecas, Aguascaliente, northern Jalisco, San Luis Potosí, Hidalgo, and probably Querétaro; marginal occurrences are: Zacatecas, Canitas and Berriozobal; Aguascaliente, 1 mi. N Chicalote; Jalisco, 9 mi. N Encarnación; Guanajuato, Celaya; Hidalgo, Irolo.
Diagnosis.—Size small (see measurements). Color dark, entire dorsal surface (h) between Cinnamon and Sayal Brown, purest on sides and flanks, upper parts strongly suffused with black; posterior surfaces and fold of pinnae of ears, white; arietiform markings, plantar surfaces of hind feet, inside of pinnae of ears, dorsal and ventral stripes of tail, blackish. Skull small; nasals long; rostrum long and narrow; interorbital region relatively wide; degree of inflation of auditory bullae about average for species; zygomatic arches light and bowed laterally; pterygoid fossae subcircular; braincase but slightly vaulted.
Comparisons.—FromDipodomys ordii ordii,D. o. palmeridiffers as follows: Size somewhat larger; color darker; skull larger; nasals longer and flaring distally; interorbital width greater; narrower across auditory bullae; interparietal region narrower; breadth across maxillary arches greater; zygomatic archesbowed laterally as opposed to relatively straight; pterygoid fossae more nearly circular; rostrum longer and narrower.
FromDipodomys ordii sennetti,D. o. palmeridiffers in larger size, darker color, longer tail and longer skull.
For comparisons withDipodomys ordii obscurusandDipodomys ordii fuscussee accounts of those subspecies.
Remarks.—Specimens from Berriozobal, Zacatecas, in the width of the rostrum and the configuration of the nasals, are intermediate betweenDipodomys ordii fuscusandD. o. palmeri. In color these animals are more nearly like typical representatives ofD. o. palmeri. The animals from Jalisco and Guanajuato are not typical but the characters differentiating them fromD. o. palmeriare merely modifications of such slight degree that they all have been placed with that subspecies.
Specimens examined.—Total, 56, distributed as follows:
Zacatecas: Canitas, 3 (USBS); Berriozobal, 10 (USBS).
San Luis Potosí: Potrero Santa Ana, 7.6 mi. S Matehuala, 2 (MVZ); Jesus Maria, 12 (USBS); 2 mi. NW San Luis Potosí, 8 (MVZ).
Aguascalientes: 1 mi. N Chicalote, 1900 m., 4 (MVZ).
Jalisco: 9 mi. N Encarnación, 1900 m., 1 (MVZ).
Guanajuato: Celaya, 8 (USBS).
Hidalgo: Ixmiquilpan, 1 (USBS); Irolo, 12 (USBS).
1. There are thirty-five recognizable subspecies of the speciesDipodomys ordiiof which four are herein, for the first time, recognized by name. Three subspecies,D. o. oklahomae,D. o. sennettiandD. o. compactus, previously were regarded by most authors as full species.2. The speciesDipodomys ordiiis divisible into six complexes, or groups, of subspecies on both geographic and morphological bases.3.Dipodomys ordiiis the most generalized Recent species of the genus.4. The extremes of geographic variation inDipodomys ordiiare greater than in any other species of the genus.5. Color, at least inDipodomys ordii, does not seem to be correlated with amount of moisture but rather with color of soil.6. Clinal variation, from north to south, is shown in the decreasing length of the nasals. This decrease in length of nasals and resultant decrease in size of the nasal chamber may be correlated with the decrease of humidity of the environment.7. Subspeciation has been enhanced by the late Quaternary mountain building which was prevalent over the western United States.
1. There are thirty-five recognizable subspecies of the speciesDipodomys ordiiof which four are herein, for the first time, recognized by name. Three subspecies,D. o. oklahomae,D. o. sennettiandD. o. compactus, previously were regarded by most authors as full species.
2. The speciesDipodomys ordiiis divisible into six complexes, or groups, of subspecies on both geographic and morphological bases.
3.Dipodomys ordiiis the most generalized Recent species of the genus.
4. The extremes of geographic variation inDipodomys ordiiare greater than in any other species of the genus.
5. Color, at least inDipodomys ordii, does not seem to be correlated with amount of moisture but rather with color of soil.
6. Clinal variation, from north to south, is shown in the decreasing length of the nasals. This decrease in length of nasals and resultant decrease in size of the nasal chamber may be correlated with the decrease of humidity of the environment.
7. Subspeciation has been enhanced by the late Quaternary mountain building which was prevalent over the western United States.
8. In general, the most primitive kinds ofDipodomys ordiioccur at the periphery of the range of the species.9. Natural selection plus geographical and ecological isolation has undoubtedly been operative in speciation and in subspeciation.10.Dipodomys desertiis found to be the most specialized species in the genus.11. Six, rather than nine, groups of species are recognized, on the basis of morphology, as comprising the genusDipodomys.12. The center of dispersal for the genusDipodomysappears to have been in the southwestern United States and the adjoining part of Mexico. A secondary center of differentiation is apparent in the low, hot valleys of central California.13. Parallel development of species is noted betweenDipodomysof the parental center andDipodomysof the isolated valleys of central California;Dipodomysin the California center is the less specialized.14. The trend of the dipodomyines, as indicated by the fossil record, has been toward a saltatorial specialization with consequent morphological changes.15. The morphological change in the direction of saltatorial specialization is clearly evident in the compacting and aligning of the viscera as well as in the lengthening of the distal segments of the hind legs and the tail, tufting of the tail, enlargement of the auditory bullae, shortening of the neck and fusion of the cervical vertebrae for stability and other modifications of the skeleton.
8. In general, the most primitive kinds ofDipodomys ordiioccur at the periphery of the range of the species.
9. Natural selection plus geographical and ecological isolation has undoubtedly been operative in speciation and in subspeciation.
10.Dipodomys desertiis found to be the most specialized species in the genus.
11. Six, rather than nine, groups of species are recognized, on the basis of morphology, as comprising the genusDipodomys.
12. The center of dispersal for the genusDipodomysappears to have been in the southwestern United States and the adjoining part of Mexico. A secondary center of differentiation is apparent in the low, hot valleys of central California.
13. Parallel development of species is noted betweenDipodomysof the parental center andDipodomysof the isolated valleys of central California;Dipodomysin the California center is the less specialized.
14. The trend of the dipodomyines, as indicated by the fossil record, has been toward a saltatorial specialization with consequent morphological changes.
15. The morphological change in the direction of saltatorial specialization is clearly evident in the compacting and aligning of the viscera as well as in the lengthening of the distal segments of the hind legs and the tail, tufting of the tail, enlargement of the auditory bullae, shortening of the neck and fusion of the cervical vertebrae for stability and other modifications of the skeleton.
Measurements (in Millimeters) of Dipodomys ordii
D. o. richardsoni1-1/2 mi. N Beaver, Oklahoma (KU)
♂ Mean (4)
251.2
131.5
40.5
41.0
24.7
22.0
4.3
14.9
13.0
26.3
Maximum
256
136
41
41.8
25.1
22.3
4.5
15.4
13.2
27.3
Minimum
243
133
40
40.4
24.4
21.5
4.2
14.6
12.7
26.0
♀ KU 17962
253
135
42
40.6
25.4
22.6
4.6
15.1
14.2
25.7
KU 17963
246
133
40
39.4
24.0
...
4.3
14.1
12.5
25.6
KU 17963
255
136
41
41.2
25.5
22.1
4.5
15.1
13.8
26.2
D. o. oklahomae2-1/4 mi. S Norman, Oklahoma (USBS)
♂ Mean (4)
239.7
130.0
39.5
39.0
23.9
21.5
4.3
13.7
13.3
24.9
Maximum
254
140
42
40.3
24.6
22.4
4.4
14.3
13.6
25.8
Minimum
227
125
37
38.0
23.0
20.9
4.2
13.4
12.8
23.9
♀ USBS 265456
245
133
40
39.1
24.5
22.0
4.4
14.4
13.5
25.3
OU 20140
234
120
39
38.7
23.8
21.7
4.4
14.3
13.5
24.8
D. o. compactus19 mi. S Port Aransas, Mustang Island, Texas (TCWC)
♂ Mean (10)
234.6
119.5
39.3
37.0
21.8
19.4
4.1
13.8
11.9
24.2
Maximum
251
134
41
37.9
22.5
20.8
4.2
14.6
12.3
25.3
Minimum
228
112
38
35.6
21.3
18.2
3.9
12.9
11.6
22.8
♀ Mean (7)
231.0
116.0
39.0
37.1
22.4
19.7
4.0
13.9
...
24.2
Maximum
235
120
41
38.2
22.9
20.5
4.1
14.7
...
24.8
Minimum
224
110
38
36.0
21.7
19.1
3.8
13.2
...
23.8
D. o. sennetti2 mi. S Riviera, Texas (TCWC)
♂ Mean (5)
217.6
112.2
35.8
37.2
23.4
20.1
4.0
13.6
13.1
24.2
Maximum
222
115
38
38.2
24.1
20.7
4.3
14.4
13.2
24.6
Minimum
208
104
34
36.3
23.0
19.4
3.8
13.0
12.6
23.8
♀ Mean (4)
218.3
112.0
36.0
37.2
23.3
20.1
4.0
13.9
13.1
24.1
Maximum
226
115
37
37.7
23.9
20.8
4.1
14.4
13.4
24.4
Minimum
209
108
35
36.7
22.8
19.0
3.9
13.3
12.6
23.5
D. o. evexusSalida, Colorado (AMNH)
♂ AMNH 28802
267.0
140.0
42.0
...
23.0
...
4.2
...
...
25.0
AMNH 28804
271
156
43
39.3
23.5
22.1
4.4
14.6
14.0
25.8
AMNH 28805
252
137
39
...
...
21.5
4.2
...
...
24.5
♀ Mean (4)
261.0
147.0
42.2
38.0
23.5
21.6
4.3
14.1
13.4
24.6
Maximum
264
149
43
38.1
24.0
22.1
4.4
14.2
13.8
25.0
Minimum
258
144
42
37.9
22.7
20.8
4.2
14.0
13.0
24.3
D. o. mediusSanta Rosa, New Mexico (USBS)
♂ Mean (6)
258.5
143.3
38.0
39.5
25.2
21.4
4.1
14.2
12.9
24.8
Maximum
266
151
40
40.6
25.9
22.5
4.2
15.2
13.1
25.6
Minimum
251
1321
37
38.9
24.8
20.6
4.0
13.5
12.8
24.1
♀ USBS 118527
261.0
145.0
40.0
40.4
25.4
22.2
4.5
15.7
13.5
25.7
USBS 127310
...
...
...
38.1
24.3
20.8
4.1
13.5
13.5
...
D. o. obscurusRio Sestin, Durango (AMNH)
♂ Mean (6)
...
...
...
36.3
22.9
19.7
3.7
13.1
12.6
22.8
Maximum
...
...
...
38.0
23.4
21.0
3.9
13.5
13.0
23.9
Minimum
...
...
...
35.2
22.6
19.1
3.6
11.9
12.4
21.7
♀ AMNH 20945
...
...
...
36.5
22.8
19.9
3.8
13.1
12.9
22.3
AMNH 20951
...
...
...
36.0
22.3
19.6
3.5
13.0
...
22.5
AMNH 20958
...
...
...
35.9
22.7
19.4
3.4
12.6
12.3
22.7
D. o. terrosusJordan, Montana
♂ AMNH 41442
280.0
155.0
44.0
42.7
26.5
23.6
4.0
15.2
14.5
27.0
AMNH 41443
267.0
155.0
40.0
40.5
24.8
21.1
4.0
14.5
13.1
25.1
AMNH 41444
279.0
162.0
41.0
40.8
25.7
21.6
4.4
14.6
13.4
25.9
♀ AMNH 41441
265.0
149.0
40.5
41.4
25.4
22.2
4.3
14.7
13.0
26.8
MVZ 25658
273.0
154.0
41.0
41.3
25.1
22.4
4.0
14.9
13.8
26.0
D. o. fremontiTorrey, Utah
♂ CM 15663
250.0
141.0
40.0
37.0
23.3
19.3
3.6
13.5
12.0
23.2
CM 15670
248.0
136.0
38.0
37.3
23.8
19.8
3.8
13.7
11.6
23.5
♀ CM 15666
258.0
141.0
39.0
37.5
23.5
19.9
3.7
13.8
11.9
23.5
CM 15667
252.0
140.0
39.0
36.8
23.5
20.5
3.7
13.5
11.5
23.2
D. o. uintensis2 mi. N Fruitland, Utah
♂ CM 11634
253.0
140.0
40.0
37.9
23.3
20.3
3.9
13.5
12.5
23.9
CM 11640
260.0
150.0
41.0
38.2
24.2
20.6
4.0
13.2
12.5
24.5
D. o. sanrafaeli12 mi. SW Green River, Utah
♂ CM 15649
265.0
144.0
41.0
...
...
...
...
14.1
...
...
♀ CM 15647
253.0
138.0
42.0
...
...
...
...
13.5
...
...
D. o. panguitchensis1 mi. S Panguitch, Utah
♂ RH 2151
257.0
145.0
41.0
...
...
...
...
13.3
12.0
...
RH 2152
252.0
135.0
40.0
...
...
...
...
13.5
11.9
...
♀ RH 2153
240.0
132.0
38.0
...
...
...
...
12.7
11.3
...
D. o. monoensis5 mi. N Benton Station, California (MVZ)
♂ MVZ 26993
232.0
125.0
40.0
36.6
23.7
20.0
3.7
13.0
...
23.4
MVZ 26995
240.0
125.0
39.0
36.8
23.0
19.8
3.6
13.4
11.5
23.3
MVZ 26997
230.0
122.0
39.0
37.2
23.2
20.1
3.6
13.4
12.0
23.9
♀ Mean (5)
228.0
125.4
38.4
36.6
23.1
19.7
3.5
13.1
11.8
23.4
Maximum
240.0
137.0
39.0
37.2
24.2
20.0
3.7
13.2
12.2
23.6
Minimum
220.0
120.0
38.0
36.0
22.6
19.3
3.4
13.0
11.5
23.0
D. o. ordiiNear El Paso, Texas (USNM)
♂ Mean (5)
...
...
...
37.1
24.1
19.6
3.7
13.0
12.7
23.5
Maximum
...
...
...
38.1
24.8
20.5
3.8
13.2
13.3
24.3
Minimum
...
...
...
36.3
23.4
18.8
3.5
12.8
12.1
22.4
♀ Mean (6)
235.2
128.0
37.4
37.3
24.0
19.6
3.7
13.3
12.8
23.5
Maximum
245.0
136.0
39.0
38.0
24.1
20.0
3.8
13.5
13.4
24.4
Minimum
222.0
120.0
35.0
36.3
24.0
19.1
3.6
13.0
12.4
23.0
D. o. luteolus1 mi. NE Casper, Wyoming (KU)
♂ Mean (12)
265.6
152.2
42.2
38.9
24.1
20.8
4.3
13.9
13.0
24.6
Maximum
281.0
163.0
43.0
39.5
25.0
22.1
4.4
14.5
13.7
25.7
Minimum
254.0
145.0
42.0
37.5
23.8
19.9
4.2
13.0
12.5
24.0
♀ Mean (7)
260.7
148.0
41.0
38.6
24.2
20.9
4.3
13.9
12.9
24.7
Maximum
269.0
153.0
43.0
40.5
25.7
21.9
4.4
14.9
13.8
25.5
Minimum
250.0
139.0
40.0
37.6
23.0
20.0
4.2
13.3
12.5
24.0
D. o. extractus1 mi. E Samalayuca, Chihuahua (MVZ)
♂ Mean (11)
235.1
128.2
37.3
37.5
24.1
20.1
3.5
13.2
13.0
23.4
Maximum
251.0
142.0
39.0
38.6
24.6
20.8
3.8
13.8
13.5
24.4
Minimum
224.0
121.0
35.0
36.2
23.2
19.3
3.3
12.6
12.4
22.3
♀ MVZ 76568
251.0
132.0
39.0
39.6
25.4
22.2
3.7
13.8
14.1
24.8
MVZ 76569
244.0
140.0
37.0
37.3
23.6
20.0
3.6
13.5
12.8
24.0
MVZ 76570
230.0
120.0
37.0
35.7
23.4
19.5
3.2
12.4
12.1
22.5
D. o. chapmaniCamp Verde, Arizona (USBS)
♂ Mean (6)
258.0
148.0
39.0
38.9
24.2
21.1
3.8
14.2
13.5
24.3
Maximum
268.0
157.0
41.0
40.1
24.7
22.1
3.9
14.8
13.9
24.7
Minimum
244.0
138.0
38.0
38.1
23.6
20.5
3.8
13.3
13.1
24.1
♀ Mean (4)
255.4
151.0
38.2
37.1
23.9
20.1
3.8
13.4
13.5
23.3
Maximum
262.0
152.0
39.0
37.7
24.3
20.1
3.9
13.7
14.9
23.5
Minimum
245.0
144.0
37.0
36.0
23.7
20.1
3.6
13.1
12.6
23.0
D. o. montanusFort Garland, Colorado (USBS)
♂ Mean (11)
252.0
140.5
41.0
37.7
24.2
20.1
4.1
13.6
12.8
23.8
Maximum
263.0
150.0
42.0
39.0
25.0
21.0
4.5
14.0
13.2
24.7
Minimum
233.0
131.0
39.0
36.6
23.6
19.0
3.9
12.7
12.4
23.0
♀ Mean (11)
256.0
141.0
40.8
37.2
23.9
19.6
4.1
13.4
12.5
23.7
Maximum
259.0
145.0
42.0
38.4
24.4
21.4
4.3
14.1
13.3
24.4
Minimum
237.0
132.0
40.0
36.0
23.0
18.2
3.9
12.3
11.9
22.6
D. o. cinderensis11 mi. SE Lund, Utah
♂ MVZ 102059
244.0
135.0
41.0
35.7
23.2
19.2
3.6
12.2
11.2
22.7
MVZ 102056
237.0
130.0
41.0
36.4
23.7
20.6
3.5
13.0
12.1
23.1
♀ MVZ 102057
229.0
122.0
41.0
36.3
23.5
20.2
3.6
13.0
11.5
23.4
MVZ 102058
241.0
127.0
40.0
36.8
24.0
20.0
3.6
13.2
11.7
23.2
D. o. fetosus2 mi. N Panaca, Nevada (MVZ)
♂ Mean (4)
242.5
133.7
42.3
37.4
23.7
20.0
3.7
13.3
11.7
23.7
Maximum
249.0
140.0
43.0
37.7
24.0
20.3
3.8
13.9
12.0
23.8
Minimum
233.0
126.0
42.0
36.8
23.2
19.5
3.7
12.7
11.5
23.6
♀ Mean (4)
229.0
125.5
40.6
35.8
23.3
19.5
3.6
12.8
11.3
22.5
Maximum
235.0
129.0
41.5
37.0
24.1
20.0
3.6
13.3
11.6
23.2
Minimum
224.0
122.0
40.0
34.3
23.0
19.0
3.6
11.9
11.0
21.9
D. o. utahensisOgden, Utah
♂ MVZ 44005
243.0
133.0
40.5
35.9
23.1
19.2
3.5
12.8
12.4
22.4
♀ MVZ 44006
238.0
133.0
38.0
36.1
...
19.8
3.7
13.5
12.2
22.9
MVZ 44007
240.0
137.0
39.0
35.6
22.9
...
3.6
12.8
11.7
22.4
MVZ 44008
241.0
134.0
40.0
36.4
23.5
19.8
3.6
13.3
12.3
...
D. o. columbianusUmatilla, Oregon
♂ MVZ 45317
243.0
140.0
40.0
37.2
23.6
20.0
3.8
13.5
12.2
23.4
MVZ 45318
236.0
137.0
40.0
36.8
23.5
20.1
3.6
13.2
...
23.4
MVZ 45322
240.0
132.0
40.0
36.4
23.4
19.9
3.7
13.0
12.1
23.3
♀ MVZ 45314
239.0
130.0
39.0
35.8
22.7
19.8
3.7
13.2
11.9
22.7
MVZ 45315
251.0
140.0
41.0
36.6
23.8
19.6
3.6
13.3
12.0
23.1
D. o. idoneus12 mi. W Lerdo, Durango
♂ UM 90027
...
...
35.0
37.3
24.3
20.7
3.8
13.0
14.1
23.1
UM 90029
230.0
129.0
35.0
37.2
23.5
20.0
3.6
13.5
13.4
23.3
D. o. priscus33 mi. S Bitter Creek, Wyoming (KU)
♂ Mean (7)
259.0
148.0
44.0
39.1
24.3
20.7
4.1
14.3
13.1
24.9
Maximum
265.0
152.0
45.0
40.4
25.1
21.2
4.3
15.2
13.6
25.5
Minimum
251.0
144.0
43.0
38.0
23.7
20.0
4.0
13.8
12.7
23.7
♀ Mean (4)
257.0
147.0
43.0
39.4
24.6
20.8
4.2
14.3
13.1
24.7
Maximum
264.0
152.0
45.0
40.4
25.2
21.9
4.3
14.9
13.3
25.2
Minimum
249.0
138.0
40.0
38.1
23.5
20.1
4.1
14.0
12.7
24.0
D. o. priscus33 mi. S Bitter Creek, Wyoming (KU)
♂ Mean (4)
208.7
109.0
40.2
35.3
22.9
19.3
3.6
12.6
11.2
...
Maximum
220.0
115.0
41.0
36.6
23.6
20.2
3.8
13.5
11.5
...
Minimum
203.0
100.0
39.0
34.0
22.1
18.4
3.5
11.7
11.1
...
♀ UU 1957
219.0
110.0
39.0
35.3
23.4
19.1
3.6
11.9
11.8
22.4
UU 1961
223.0
120.0
40.0
37.0
23.6
20.4
3.6
13.1
12.1
23.9
D. o. cineraceusDolphin Island, Great Salt Lake, Utah
♀ USNM 263893
228.0
129.0
39.0
37.2
23.4
19.7
3.5
13.6
11.6
23.0
USNM 263894
230.0
132.0
38.0
37.1
23.7
20.1
3.6
13.4
11.9
23.6
D. o. marshalliStansbury Island, Great Salt Lake, Utah
♂ UU 2968
238.0
128.0
40.0
36.4
23.5
...
3.5
13.0
12.0
23.2
UU 2969
241.0
136.0
40.0
36.3
23.7
19.2
3.7
12.9
...
23.0
♀ UU 2972
...
...
...
35.9
23.5
19.9
3.6
12.9
11.9
23.1
D. o. inaquosus11 mi. E and 1 mi. N Jungo, Nevada (MVZ)
♂ Mean (4)
243.0
137.0
40.0
36.6
23.5
19.8
3.7
13.5
11.8
23.1
Maximum
247.0
140.0
40.5
36.8
24.0
20.0
3.8
13.8
12.1
23.5
Maximum
238.0
131.0
39.0
36.4
23.3
19.5
3.6
13.1
11.5
23.0
♀ MVZ 73577
242.0
139.0
40.5
36.4
23.9
19.2
3.7
13.2
11.4
22.9
MVZ 73579
231.0
130.0
40.5
36.3
23.2
19.0
3.5
12.7
11.0
23.2
MVZ 73582
254.0
138.0
41.0
38.2
23.9
20.0
3.6
13.9
12.2
...
D. o. attenuatusJohnson's Ranch, Texas
♂ TCWC 3633
235.0
126.0
41.0
37.3
24.1
20.1
3.7
13.7
...
23.4
TCWC 3634
237.0
136.0
37.0
35.5
22.1
18.8
3.5
13.2
12.2
22.3
UM 79121
245.0
143.0
37.3
35.7
22.3
19.1
3.6
12.4
12.9
23.3
♀ UM 79122
238.0
140.0
38.0
34.6
22.5
18.5
3.5
11.9
13.0
21.9
D. o. fuscusJuamave, Tamaulipas (USBS)
♂ Mean (4)
248.0
149.0
38.5
37.6
24.1
20.2
3.6
12.6
12.7
23.6
Maximum
254.0
152.0
40.0
38.3
24.7
20.4
3.7
13.1
12.9
23.7
Minimum
241.0
146.0
38.0
37.0
23.5
19.7
3.6
12.3
12.5
23.6
♀ Mean (4)
244.7
146.2
38.5
37.4
24.1
19.9
3.4
12.9
12.7
23.2
Maximum
254.0
155.0
40.0
37.7
24.5
20.5
3.6
13.0
12.9
23.4
Minimum
240.0
138.0
38.0
37.2
23.7
19.2
3.3
12.7
12.4
23.2
D. o. longipesKayenta, Arizona
♂ USBS 247915
255.0
145.0
41.0
39.3
25.5
20.7
3.9
13.8
13.0
24.3
USBS 247916
242.0
138.0
40.0
38.4
25.4
20.6
3.8
13.3
12.7
24.6
♀ USBS 247552
255.0
135.0
40.0
39.9
25.7
21.2
4.0
13.8
...
25.4
D. o. pallidus18 mi. SW Orr's Ranch, Utah (UU)
♂ Mean (4)
236.6
131.3
42.3
37.6
24.2
20.4
3.8
13.4
11.8
23.6
Maximum
240.0
134.0
43.0
37.9
24.6
20.9
4.0
13.6
12.1
24.2
Minimum
230.0
128.0
41.0
37.0
23.7
19.8
3.7
12.9
11.5
23.4
♀ UU 3528
229.0
132.0
40.0
36.6
23.5
20.0
3.7
12.5
12.0
...
D. o. nexilis5 mi. W Naturita, Colorado
♂ USBS 149940
261.0
156.0
44.0
38.7
25.1
18.9
4.1
13.8
11.4
24.6
♀ USBS 149941
265.0
142.0
45.0
40.5
26.3
21.5
4.2
14.4
13.1
25.3
D. o. cupidineusHouserock Valley, Arizona (USBS)
♂ Mean (6)
239.0
138.0
39.8
36.9
24.1
19.3
3.8
12.8
12.0
23.6
Maximum
244.0
146.0
41.0
37.5
24.9
20.1
4.0
13.0
12.6
24.3
Minimum
225.0
130.0
38.5
36.3
23.5
18.7
3.6
12.6
11.7
23.2
♀ Mean (4)
244.7
141.0
40.0
37.4
24.2
19.7
3.8
13.2
12.3
23.9
Maximum
247.0
147.0
40.5
37.7
24.4
20.0
3.9
13.5
12.6
24.6
Minimum
240.0
137.0
39.5
37.1
23.9
19.5
3.7
12.9
12.0
23.7
D. o. palmeriJesus María, San Luis Potosí (USBS)
♂ Mean (6)
242.0
141.6
37.6
36.7
23.6
20.0
3.3
12.9
12.9
22.6
Maximum
255.0
157.0
39.0
37.3
24.4
20.8
3.6
13.6
13.4
23.0
Minimum
229.0
129.0
35.0
35.9
23.2
19.4
3.0
12.5
12.6
22.0
♀ Mean (6)
239.5
139.6
37.1
36.4
23.8
19.9
3.4
12.8
13.1
22.7
Maximum
252.0
148.0
38.0
37.3
23.9
20.0
3.6
13.3
13.2
23.5
Minimum
225.0
128.0
36.0
35.9
23.7
19.5
3.3
12.2
13.0
22.0