Larger ImagePl.92.Scotch Large Heath. 1, 2, 5male; 3, 4female.Small Heath. 6, 9, 10, 14male; 7, 8, 11, 12, 13female.
Pl.92.Scotch Large Heath. 1, 2, 5male; 3, 4female.Small Heath. 6, 9, 10, 14male; 7, 8, 11, 12, 13female.
Pl.92.
Scotch Large Heath. 1, 2, 5male; 3, 4female.
Small Heath. 6, 9, 10, 14male; 7, 8, 11, 12, 13female.
Pl.93.Small Heath.Eggs, natural size and enlarged; caterpillar and chrysalis.
Pl.93.Small Heath.Eggs, natural size and enlarged; caterpillar and chrysalis.
Pl.93.
Small Heath.
Eggs, natural size and enlarged; caterpillar and chrysalis.
The life-history is figured on Plate95—the lower set of figures.
The egg is described by Newman "as a depressed sphere and white," and he states, "it is attached to the twigs of blackthorn (Prunus spinosa) in the autumn, often as late as the end of September or beginning of October; it is not hatched until the spring."
The caterpillar is bright pale green, and the lines on the back and sides are yellowish, as also are the oblique streaks on the sides and the border of the ridge above the feet. There are some bristles along the ridge on the back and also on that above the feet. It feeds on blackthorn in May and June, and will eat the foliage of almost any kind of plum. I have reared fine specimens from caterpillars which fed on greengage.
The chrysalis is pale reddish-brown with a dark line downthe middle of the back and some pale oblique streaks on each side; the wing-cases are freckled with darker brown. Barrett, quoting Fenn, says, "Suspended by the tail and a silken girth to the stem of the food-plant close to the ground." Those that I have seen pupated on or under leaves, and so far as I could observe without any girth, and certainly not suspended.
Nearly two hundred years ago the male of this butterfly was known as the Brown Hairstreak, whilst the female was called the Golden Hairstreak. The caterpillar seems to have been observed in quite early times. It has always been a local species, and although it appears to frequent hedgerows occasionally, its haunts generally are open grounds in the neighbourhood of woods, where blackthorn or sloe is plentiful. August and September are the months for the butterfly, but it does not seem to be very often observed on the wing, even in places where the caterpillars are known to occur. When seen it is generally high up on, or around, some oak tree. Occasionally, however, it visits the bramble blossoms, and at such times becomes a fairly easy prey. The caterpillar is obtained by beating sloe bushes.
Barrett, who seems to have worked out its distribution in England and Wales pretty closely, remarks, "In the eastern counties it has been taken occasionally in Norfolk and Suffolk, more frequently in Essex, where, in Epping Forest, it has been fairly common; also in Cambridgeshire, Huntingdonshire, and Northamptonshire, in some plenty. In very few localities in Kent, Sussex, Hants, and Dorset; rarely in Gloucestershire, and possibly Somerset; but found in many Devonshire localities, especially in the sheltered valleys around the Dartmoor range, and in the charmingly wooded districts about Axminster and Sidmouth; becoming common towards Dartmouth. It has also been found commonly near Marlborough, Wilts, and plentifully in some parts of North Wales; apparently rare in South Wales, but certainly existing in some parts of thewooded districts skirting Milford Haven. Also recorded from Worcestershire, and Cannock Chase in Staffordshire; and northward in the favoured districts of Grange and Silverdale in North Lancashire, and Witherslack in Westmoreland." As Surrey is not quoted in the foregoing, it may be mentioned as one of the counties in which the species is found. In Ireland Kane says that it is "abundant in certain localities in Munster; and in Co. Galway at Claring Bridge, and Oranmore; Cork; Killoghrum Wood, Enniscorthy; Blarney, Killarney."
Larger ImagePl.94.Brown Hairstreak. 1,male; 2, 3female.
Pl.94.Brown Hairstreak. 1,male; 2, 3female.
Pl.94.
Brown Hairstreak. 1,male; 2, 3female.
White-letter Hairstreak. 4, 6male; 5, 7female.
White-letter Hairstreak. 4, 6male; 5, 7female.
Pl.95.White-letter Hairstreak.Egg enlarged; caterpillar and chrysalis.Brown Hairstreak.Egg enlarged; caterpillar and chrysalis.
Pl.95.White-letter Hairstreak.Egg enlarged; caterpillar and chrysalis.Brown Hairstreak.Egg enlarged; caterpillar and chrysalis.
Pl.95.
White-letter Hairstreak.Egg enlarged; caterpillar and chrysalis.
Brown Hairstreak.Egg enlarged; caterpillar and chrysalis.
It is distributed throughout Central and Northern Europe, except the Polar region, and its range extends through Northern Asia to Amurland, Ussuri, and China.
The butterfly (figured on Plate96) has the sexes differently ornamented, as in the last species. The male is strongly tinged with purplish-blue, the veins are blackish; the outer margin of the fore wings are narrowly, and the costa and outer margin of the hind wings are broadly, bordered with black. The female is purplish-black, with two patches of bluish-purple in the discal cell and space below; often there is a smaller patch of the same colour between them, the whole forming a large blotch interrupted by the blackish veins. Under side greyish with blackish shaded white lines; two or three blackish clouds on the outer margin of fore wings above the inner angle; these are sometimes edged with orange; a black spot on anal angle of the hind wings, with an orange one above it, and a black-centred orange spot between veins 2 and 3.
Variation in this species is exceptional. An aberration known asbella, Gerhard, has a yellowish mark at end of the cell on the upper side of the fore wings, and at least one such variety has been taken in England. Sometimes the blotch on the female is rather blue than purple; a male specimen with blue streaks on the costa of the fore wings has been recorded, andBarrett mentions a gynandrous specimen in which the right side was that of the male.
The egg is pale brown tinged with pink, and over this is a whitish network. The caterpillar is reddish-brown and downy; a black line along the back has a whitish edge, and there are whitish oblique stripes, with blackish edge, on each side of the central line; the segmental divisions are well marked, and the spiracles are blackish with pale rings. The head, which, when the caterpillar is resting, is hidden within the first body ring, is brownish and glossy, and there is a greyish shield-like mark on the second ring. The chrysalis is red-brown, with darker freckles; the body is downy, and there are traces of oblique marks thereon. It does not appear to be fastened by the tail, but the cast larval skin remains attached; there are a few strands of silk around and about the chrysalis, but these are very flimsy, although they hold it in position on the ground or under a leaf.
The eggs are laid in July or August on twigs of oak, but the caterpillars, it is said, do not hatch out until the following spring. In May and early June the caterpillars are full grown, and may be obtained by beating or jarring the branches of oak trees in places where the butterfly is known to occur. They have also been found on sallow.
This species frequents oak woods, or the borders thereof, in July and August, and is often more easy to see than to capture, as it has a tantalizing trick of flying around the upper branches of the trees. Occasionally it resorts to lower growing aspens, probably to feast on the honey dew, the secretions of Aphides, with which the leaves are often covered in hot summers. It seems to be pretty generally distributed in all parts of England and Wales, and in Scotland as far north as Ross. In Ireland it appears to be more local, and has only been recorded from the east and south.
It is found in all parts of Europe, except the northern.
Larger ImagePl.96.Purple Hairstreak. 1male; 2, 3female.Black Hairstreak. 4, 6male; 5female.Green Hairstreak. 7male; 8, 9female.
Pl.96.Purple Hairstreak. 1male; 2, 3female.Black Hairstreak. 4, 6male; 5female.Green Hairstreak. 7male; 8, 9female.
Pl.96.
Purple Hairstreak. 1male; 2, 3female.
Black Hairstreak. 4, 6male; 5female.
Green Hairstreak. 7male; 8, 9female.
Black Hairstreak.Eggs, natural size and enlarged; caterpillar and chrysalis.
Black Hairstreak.Eggs, natural size and enlarged; caterpillar and chrysalis.
Black Hairstreak.Eggs, natural size and enlarged; caterpillar and chrysalis.
Purple Hairstreak.Egg enlarged; caterpillar and chrysalis.
Purple Hairstreak.Egg enlarged; caterpillar and chrysalis.
Purple Hairstreak.Egg enlarged; caterpillar and chrysalis.
Pl.97.Green Hairstreak.Eggs, natural size and enlarged; caterpillar and chrysalis.
Pl.97.Green Hairstreak.Eggs, natural size and enlarged; caterpillar and chrysalis.
Pl.97.
Green Hairstreak.Eggs, natural size and enlarged; caterpillar and chrysalis.
This butterfly is figured on Plate96.In colour it is dark brown or, when quite fresh, brownish-black; there are some orange marks on the outer margin of the hind wings, and these are most distinct in the female, in which sex there are orange spots on the fore wings also. The male has a pale sexual mark at the end of the cell of the fore wings, but this is less distinct than in the following species. The under side is brown, with a bluish-white interrupted transverse line on each wing, that on the hind wings angled before reaching the inner margin. All the wings have an orange band on the outer margin, but on the fore wings of the male this is often indistinct; there are some white-edged black spots before it, and, on the hind wings, beyond it also.
The eggs are laid in July on the twigs of blackthorn, but the caterpillars do not hatch until the following spring. The egg figured on Plate97was reddish-brown and appeared rather shiny. The caterpillar is described as yellowish-green, with a darker green furrow and purplish ridges along the back; the latter are edged with whitish and the divisions between the rings are yellowish. The head is pale brown. The chrysalis, which is attached by the tail and has a silken thread around it, is black, marked on the head and body with yellowish-white.
The caterpillars feed on blackthorn (Prunus spinosa) in a state of nature, but will eat the leaves of damson in confinement. They may be obtained in May, in their particular haunts, by beating sloe bushes with a beating tray, or an inverted open umbrella, held under to intercept the evicted caterpillars, etc.
This butterfly was not known as British until 1828, when a number of specimens were captured at Monkswood in Huntingdonshire. These were sold by the captor asT. w-album, which was then called the Black Hairstreak. As soon as the mistake was detected, it was given out that thespecimens had been taken in Yorkshire, but this was only a ruse, asT. prunihas never occurred in that county. It is confined, so far as Britain is concerned, to three or four of the midland counties. "Mr. Herbert Goss, who has found it at Barnwell Wold, and in other wooded districts of Northamptonshire, at intervals, for more than twenty years past, says that it is fond of sitting on the flowers of privet (Ligustrum), and ofViburnum lantana, in the woods, and sometimes is to be found in numbers. Its time of emergence is very variable, apparently regulated by the lateness of the spring—from June 17th to the first week in July. Reared specimens made their appearance from June 13th to 27th. He writes, 'It was the greatest possible pleasure to see them walking about the table while I was at breakfast.' In 1858 it was found commonly at Kettering, and in 1859 at Oundle, and has been recorded at Warboys Wood, Huntingdonshire, and in Buckinghamshire. One specimen was taken at Brandeston, Suffolk, by the Rev. Joseph Green; and Mr. Allis found it commonly in the Overton Woods and about St. Ives. There is also a record in Monmouthshire, which may require confirmation. This butterfly does not appear to be losing ground in this country, its fondness for trees and lofty bushes rendering it difficult to capture" (Barrett).
A writer in theEntomologistfor 1874 mentions Linford Woods, in Bucks, as a locality where he had observed several specimens, mostly females, on flowers of privet.
It is found throughout the greater part of Europe and also in Amurland and Corea.
The male of this butterfly (Plate94) is blackish, with a small whitish sex mark at end of the discal cell of the fore wing; there is a small orange spot at the anal angle of the hind wings. The female agrees in colourwith the male, but the tails are longer, and there is no sex mark on the fore wings. The under side is brownish, with a white line on each wing, that on the hind wings forming aWbefore the inner margin; the hind wings have a black-edged orange band on the outer margin which is finely tapered towards the costa. Captured specimens are usually browner than those that are reared from caterpillars.
The species does not exhibit much tendency to variation. The white lines on the under side may be rather broad or very narrow, and that on the hind wings is sometimes so broken up towards the inner margin that theWcharacter disappears; when absence of the anal orange spots on the upper side is associated with the broken line, the form is known asbutlerowi. I have several males without theW, and some of these have the orange spot above, whilst others are without it. Barrett refers to a specimen in which there is "on the under side an extension of white colour from the white line towards the margin, in the fore wings forming a broad wedge-shaped band, but in the hind wings occupying the whole space from the white line to the orange band."
The egg has been described as whitish in colour, and in shape something like an orange with a depression on the top. The eggs are laid on twigs of elm in July, and, according to some writers, remain thereon throughout the winter. The caterpillar when full grown is yellowish-green and covered with short hairs; the ridges on the back are yellowish, and there are oblique whitish streaks on each side of the darker dorsal line. The head is black. When about ready to assume the chrysalis state, the whole body becomes purplish-brown. The chrysalis is brownish, sometimes tinged with purple; covered with tiny bristles except on the blackish wing cases, and there are two purplish lines on the back. It is attached by the tail, and has a strand or two of silk around it, generally on the under side of a leaf.
In a state of nature the caterpillar feeds on wych-elm (Ulmus montana), but it will eat the leaves of the common elm (Ulmus campestris). It is to be obtained in May and June by beating wych-elms in localities where the butterfly is known to occur.
The butterfly is on the wing in July, and usually disports itself around the elm trees, but it is fond of bramble blossoms, and may often be netted when feasting on those flowers. It is a local species, but, as a rule, plentiful enough in its localities. It is rare in Hampshire and Dorsetshire, scarce in Sussex, and not found in many parts of Kent. Ripley, in Surrey, was a well-known locality for it in the early part of the last century, and the caterpillars were found there commonly quite recently. In Essex it is generally common near Maldon. And, according to Barrett, it is "plentiful in various parts of Suffolk; very scarce in Norfolk; found more or less plentifully in Herts, Hants., Cambs., and Northamptonshire; very rare in Nottinghamshire; but again to be found in North Lincolnshire; and common in several localities near Doncaster, Barnsley, and elsewhere in Yorkshire. This appears to be its northern limit, and in this respect it contrasts curiously withThecla betulæ[The Brown Hairstreak], since it extends farther north in the east than that species; yet in the west is recorded no farther than Cheshire and Shropshire, where I found it thirty-five years ago upon Benthall Edge. In Herefordshire it is recorded but rarely; more commonly in Worcestershire; also in Derbyshire and Needwood Forest, Staffordshire; common around Burton-on-Trent and elsewhere in Leicestershire; and in Oxfordshire, Bucks, and Berks. But its metropolis seems to be Wiltshire, where Mr. Perkins has found it around Marlborough and Savernake in thousands, as well as in Gloucestershire." It has also been obtained in Monmouthshire, but its extreme western limit seems to be Weston-super-Mare, Somersetshire. Abroad it is widely distributed in Europe, except theextreme north and south-west; its range extends into Asia Minor, and to Amurland and Japan.
Larger ImagePl.98.Large Copper.1, 4,male; 2, 3, 5female.
Pl.98.Large Copper.1, 4,male; 2, 3, 5female.
Pl.98.
Large Copper.
1, 4,male; 2, 3, 5female.
Pl.99.Large Copper.Caterpillar and chrysalis.
Pl.99.Large Copper.Caterpillar and chrysalis.
Pl.99.
Large Copper.
Caterpillar and chrysalis.
Thecla spiniandT. ilicis, two species of Hairstreak butterflies belonging to Central and Southern Europe, have been mentioned as occurring in Britain by some of the earlier authors. There is not, however, the slightest reason to suppose that either of them ever occurred naturally in this country.
Both sexes of this butterfly (Plate96) are brown with a faint golden tinge above, and green on the under side. The male has a dark, or, when the plumules are dislodged, pale sexual mark, which is oval in shape, and placed at the upper corner of the discal cell in the fore wings. Occasionally there are some orange scales at the anal angle of the hind wings, and more rarely, and in the female, at the extremities of veins two and three also. On the under side of some specimens, chiefly from Northern localities, there is a transverse series of white dots across all the wings; more often these are confined to the hind wings, and sometimes they are almost or quite absent from all the wings. Now and then the under side of the hind wings is found to be brown in colour, and this change in colour has been ascribed to the action of moisture. The life-history is figured on Plate97.
The egg is greenish, reticulated with paler or with whitish-green; the reticulation is somewhat rough on the side, but becomes finer towards and on the top, which has the centre hollowed. Laid on the petals of the common furze (Ulex europæus), and on leaves of rock-rose (Helianthemum chamæcistus).
The caterpillar feeds in June and July. It is pale green, with a darker line along the back, and yellow oblique stripes on the sides. Among the plants that it has been found upon, or is known to eat, are dyer's greenweed (Genista tinctoria), needlefurze (G. anglica), broom (Cytisus scoparius), dwarf furze (Ulex nanus), whortleberry (Vaccinium myrtillus); also the berries of buckthorn (Rhamnus), making holes through which the contents of the berry is extracted; buds of bramble (Rubus), and of dogwood (Cornus sanguinea), are also attacked in a similar way.
The chrysalis is clothed with tiny hairs, and when freshly formed is green in colour, but becomes purplish-brown after a time. It appears to be unattached to anything. I think, however, that there are generally a few strands of silk around or about it, but these are so easily broken when the chrysalids are removed that they escape observation. May and June are the months for the butterfly, which occurs in various kinds of situations, such as the outskirts of woods, high hedgerows, hill slopes, and boggy heaths. I once saw it in abundance about the entrance from Lynton to the Valley of Rocks. Its resemblance on the under side to the leaves on which it perches is as baffling to the collector as is the resting habit of the Grayling butterfly previously referred to. It seems to be pretty generally distributed throughout the kingdom, but is rather more local in Ireland than elsewhere, and it has not yet been recorded from the Orkney or Shetland Isles. Its range extends throughout the Palæarctic Region.
The brilliant butterfly, figured on Plate99, is of a coppery orange colour. In the male the fore wings have two black dots in the discal cell, the outer one linear, and the outer margin is narrowly blackish; the hind wings have a linear black mark in the cell, and the outer margin is narrowly edged with blackish and dotted with black. The female is more conspicuously marked with black; there are two, sometimes three, spots in the cell of the fore wings, and a transverse series of sevenor eight beyond; the outer margin is broadly bordered with black, and there are generally two spots above the inner angle; the hind wings have a black spot in the cell, and a series of black spots beyond, but the whole basal three-fourths of these wings is often deeply suffused with blackish; the outer margin is bordered and spotted with black. The sexes are much alike on the under side, and have reddish-orange fore wings with bluish grey outer margins, and black spots as on the upper side of the female; the hind wings are bluish-grey, powdered with bluish towards the base, and with whitish ringed black spots; five of these spots are before the linear discal mark, and a series of nine or ten beyond; an orange band on the outer margin has black dots on each edge.
Except as regards the size and the shape of the spots, especially in the female, there appears to have been but little variation noted in this species in England.
The two fine female specimens figured on the plate have a more or less distinct wedge-shaped black spot in the basal end of the discal cell of the fore wings. Dale mentions that he has an "almost entirely black" example of the female in his collection.
The var.rutilus, which is the continental form of our butterfly, is smaller in size, as a rule, the spots are not so large, and the orange band is always narrower on the under side of the hind wings. It has been averred that some of the British specimens are referable to this form.
Newman, writing about 1870, gave the following life-history details:—"The egg is laid on the leaves of the great water-dock (Rumex hydrolapathum) during the month of August, and the young caterpillars (never, to the best of my belief, observed) probably emerge during the following month, and hibernate very early at the base of the petioles.
"The caterpillar is full fed in June, and then lies flat on the dock-leaf, rarely moving from place to place, and, when it doesso, gliding with a slug-like motion, the legs and claspers being entirely concealed. The head is extremely small, and can be completely withdrawn into the second segment: the body has the dorsal surface convex, the ventral surface flat; the divisions of the segments are distinctly marked, the posterior margin of each slightly overlapping the anterior margin of the next, and the entire caterpillar having very much the appearance of aChiton; the sides are slightly dilated, the legs and claspers are seated in closely approximate pairs, nearly on a medio-ventral line. The colour is green, scarcely distinguishable from that of the dock-leaf; there is an obscure medio-dorsal stripe, slightly darker than the disk, and in all probability due to the presence of food in the alimentary canal. The chrysalis is obese, blunt at both extremities, attached by minute hooks at the caudal extremities, and also by a belt round the waist." Newman adds, "My acquaintance with the caterpillar and chrysalis was made very many years ago in Mr. Doubleday's garden at Epping, where the very plant ofRumex hydrolapathum, on which the caterpillars fed, is still in existence."
The caterpillar was described by Stephens, in 1828, as somewhat hairy, bright green, with innumerable white dots. The same author states that the chrysalis was "first green, then pale ash-coloured, with a dark dorsal line and two abbreviated white ones on each side, and, lastly, sometimes deep brown."
The figure of the caterpillar on Plate98is after Westwood, and that of the chrysalis after Newman ("Grammar of Entomology").
Although he refers to it as "hippothoë," the Large Copper seems to have been known to Lewin (1795), as he states that specimens had been taken in Huntingdonshire. Haworth (1803) mentions its occurrence in the fens of Cambridgeshire, and Stephens, twenty-five years later, wrote:—"This splendid insect appears to be confined to the fenny counties of Cambridge and Huntingdon, with the neighbouring ones of Suffolkand Norfolk, unless the account of its capture in Wales by Hudson be admitted; but this may probably be the following species[hippothoë], which may, moreover, eventually prove synonymous withLy. dispar. In the first two localities it appears to occur in great profusion, as several hundred specimens have been captured within these last ten years by the London collectors, who have visited Whittlesea and Yaxley Meres, during the month of July, for the sole purpose of obtaining specimens of this insect."
Pl.100.Small Copper.Egg, natural size and enlarged; caterpillars and chrysalids.
Pl.100.Small Copper.Egg, natural size and enlarged; caterpillars and chrysalids.
Pl.100.
Small Copper.
Egg, natural size and enlarged; caterpillars and chrysalids.
Larger ImagePl.101.Small Copper.1, 2Typical male; 3typical female; 4, 5, 6, 8, 9, 10, 11, 12varieties; 7var.schmidtii.
Pl.101.Small Copper.1, 2Typical male; 3typical female; 4, 5, 6, 8, 9, 10, 11, 12varieties; 7var.schmidtii.
Pl.101.
Small Copper.
1, 2Typical male; 3typical female; 4, 5, 6, 8, 9, 10, 11, 12
varieties; 7var.schmidtii.
Dale states that "the latest capture, consisting of five specimens, appears to have been made in Holme Fen, by Mr. Stretton either in 1847 or 1848."
There is evidence that floods, which were not uncommon in the home of the Large Copper, were not really injurious to the butterfly, and therefore the occasional submergence of its feeding grounds can hardly have been the cause of its almost sudden destruction. It seems more probable that its disappearance was due to the draining of the fens, and at least it is significant that the two events were almost coincident.
There are records of the butterfly having been taken in various odd localities since it was last seen in fenland, but the latest of those dates back to the year 1865. There seems to be no question that the butterfly is now extinct in England, and, lamentable to relate, the chief locality where we can hope to secure a specimen or two for our collection is in the neighbourhood of Covent Garden, where the only requirement for the capture will be a well-lined purse.
The continental formrutilusis found in Germany, France, Northern Italy, South-Eastern Europe (except Greece), Northern Asia Minor, Armenia, and the Altai. The Asian formauratusoccurs in South-Eastern Siberia, Amurland, Corea, Northern China, and Amdo.
Four other kinds of "Coppers" have been reported asoccurring in England: these areChrysophanus hippothoëandC. virgaureæ, both of which have even had English names bestowed upon them, to wit, the Purple-edged Copper and the Scarce Copper;C. gordius, andC. circe(dorilis). These are only mentioned to afford an opportunity for saying that there does not appear to be the least reason for considering either of them to be a British butterfly. Kirby, Barrett, and others, however, think it possible that the first two may have inhabited England in ancient times.
This little butterfly is very smart, in activity as well as appearance. In colour it is very similar to the last species, but both sexes are spotted with black on the fore wings, the outer series of six spots forming a very irregular row; the hind wings are black, with a wavy orange-red band on the outer margin.
There is considerable variation, and it is, therefore, deemed advisable to give a number of figures representing some of the more striking aberrations. The three figures at the top of Plate101depict the normal male and female; the latter sex is Fig.3.For the loan of the other specimens (Figs. 4-12) my thanks are due to Mr. E. Sabine, who has a very fine and extensive series of varieties of this butterfly. Other examples of aberration on the under side are shown on Plate119.Blue spots are sometimes found on the hind wings; these are placed near the orange-red band, and occasionally they attain a good size. Specimens much suffused with blackish sometimes occur; these are referable to var.eleus, which is the usual summer form in some of the warmer countries abroad. A very rare form is that known asschmidtii(Fig.7), in which the ground colour of the fore wings and the band on the hind wings are silvery white instead of orange or coppery-red. A modificationof this form which is hardly less rare has a creamy tint. Straw-coloured or pale golden specimens are rather more frequently met with. The colour of the hind wings in fresh specimens is sometimes steely-grey, but blackish is the more usual hue; the band on the outer area, which as a rule agrees in colour with the fore wings, varies in width a good deal, and occasionally is more or less obscured by the blackish ground colour. The arrangement, size, and shape of the black spots, both above and below, are subject to much vagary, sometimes of a very striking kind, as, for example, when the spots of the outer series on the fore wings are united with the discal pair and form a large irregular blotch. A remarkable specimen taken some years ago in the Isle of Wight had a small patch of copper with a black spot in it on the under side. This gave one the idea of a clumsy attempt at patching, but as I happened to take that particular specimen, I know that it had not been tampered with. Gynandrous specimens of this butterfly sometimes occur, but these are very rare.
The egg is of a yellowish-white colour at first, and afterwards becomes greyish; the pattern on the shell, which resembles network, is always whiter.
The caterpillar is green and similar in tint to the leaf of dock or sorrel upon which it feeds. It is clothed with short greyish hair which arises from white dots; the dorsal line is brownish-olive, and the ring divisions, especially along the back, are well defined. Head very small, pale brownish, marked with blackish, drawn into the first ring of the body when resting. The legs and prolegs are tinged with pink, and sometimes the body is marked with pink.
The chrysalis is pale brown, sometimes tinged with greenish, and freckled with darker brown; there is a dark line along the middle of the thorax and body, the wing cases are streaked with blackish, and the body is dotted with black. Attached by the tail and loose silken threads around the body to a leaf or stem.
There seem to be three broods of this species in most years: the first is on the wing in May, sometimes in April; the second in July or early August; and the third in early October. It is not a difficult species to rear from the egg, and as varieties appear to be most frequent in the third brood, the eggs should be obtained from females of the second brood. Dock and sorrel (Rumex) are the food-plants of the caterpillar, and these are most useful in a growing condition.
The butterfly frequents all kinds of open situations, and is fond of basking upon flowers, more particularly those of the Compositæ, from which vantage ground it dashes with great alertness at any other small butterfly that may happen to fly that way. Whether these seeming attacks are really due to pugnacity, as has been stated by some writers, or are merely of a sportive character, is not altogether clear. As, however, the meeting of the two butterflies usually results, when both are Small Coppers, in a series of aërial evolutions by the pair, it would seem that there is a good deal of playfulness in the business. After the gambol is over, one butterfly may dart off with the other in hot pursuit, and then both move so rapidly that their course is difficult to follow. If the butterfly intercepted happens to be a Blue or a Small Heath, the Copper returns to the flower from which it started, and prepares for another raid when the opportunity offers. It occurs throughout the United Kingdom, but in Scotland it does not extend northwards beyond the Caledonian Canal.
Abroad it is found throughout the Palæarctic Region, and is represented in North America by the formhypophlæas.
The male is purplish-blue suffused with fuscous, especially on all margins except the inner one; there are two velvety black spots encircled with pale blue at the anal angle of the hindwings, and a slender black tail, tipped with white, appears to be a continuation of vein 2. The under side is grey-brown, with numerous white wavy lines and broader streaks; there is a whitish band on each wing before the outer margin, and black spots as above, but these are ringed with metallic blue.
Short-tailed Blue.Eggs enlarged.
Short-tailed Blue.Eggs enlarged.
Short-tailed Blue.Eggs enlarged.
Pl.102.Long-tailed Blue.Caterpillar and chrysalis (after Millière).
Pl.102.Long-tailed Blue.Caterpillar and chrysalis (after Millière).
Pl.102.
Long-tailed Blue.Caterpillar and chrysalis (after Millière).
Larger ImagePl.103.Long-tailed Blue.1male; 2, 3female.Short-tailed Blue.4, 6male; 5, 7female.
Pl.103.Long-tailed Blue.1male; 2, 3female.Short-tailed Blue.4, 6male; 5, 7female.
Pl.103.
Long-tailed Blue.
1male; 2, 3female.
Short-tailed Blue.
4, 6male; 5, 7female.
I have not seen any of the early stages of this butterfly. The caterpillar, which feeds upon the green seeds in pods of the Leguminosæ, including the garden pea and the lupine, is figured on Plate102.It is described as being green or reddish-brown in colour, with a dark stripe on the back, double oblique lines on the sides, and a white line below the yellow spiracles; head black. The chrysalis is of a red or yellowish colour, and dotted with brown. It has a silken girdle and is said to be attached to a stem, as shown in the figure, but probably it is more often fixed up among the withered leaves of the food-plant. Two of the earliest known British specimens of this butterfly were taken by the late Mr. Neil McArthur on August 4th and 5th, 1859, on the Downs at Brighton; the third example was captured by Captain de Latour at Christchurch, where it was flying about a plant of the everlasting pea in his garden on August 4th of the same year. Newman has noted that in that particular year the butterfly was very abundant in the Channel Islands and on the coast of France. No other specimen seems to have been observed in England until 1879, in which year one was taken at Freshwater in the Isle of Wight on August 23rd. In 1880 a specimen was captured in a garden near Bognor, Sussex, on September 12th. On October 2nd, 1882, one was obtained at West Bournemouth. Three were netted in 1893, one of these in late August, and one in the third week of September, both in Sussex; the third was taken in Kent (inland) in September. In 1899 a specimen was found at Winchester on September 1st, and one at Deal on the 16th of the same month; each of these, curiously, was sitting on a window. On August 2nd, 1904, one example was taken in agarden near Truro, Cornwall. In addition to the above, single specimens have been reported as taken at Brighton, July, 1890, and at Heswell, Cheshire, in 1886 or 1887.
It will thus be seen that the occurrence of this butterfly in England is exceedingly infrequent. The species is common in Africa and in Southern Europe; thence it extends eastward through Asia to China and Japan, and southwards to Australia. It is also found in the Sandwich Islands. It is believed to be migratory in its habits, and it is supposed that the occasional specimens that arrive in this country come to usviâthe west coast of Europe.
In its proper home there is a succession of broods of the butterfly, and if by chance a few females were to visit this country in the early summer, they most probably would lay eggs, and the caterpillars resulting from these would almost certainly be able to feed up and attain the perfect state here. So far there is no reason to suppose that the caterpillar has ever occurred in England.
The interesting little butterfly represented on Plate103was not known to occur in Britain until 1885, when the Rev. O. Pickard Cambridge made the startling announcement that his sons had captured two specimens, a female on August 18th, and a male on August 20th of that year, the scene of capture being Bloxworth Heath, Dorset. Shortly after this fact was made public the Rev. J.S. St. John added a record of two males that he had discovered in a small collection of Lepidoptera made by Dr. Marsh, who stated that he had taken the specimens ofC. argiadesin 1874, close to a small quarry near Frome. In addition to these a specimen, also recorded by Mr. Cambridge, was taken at Bournemouth in August, 1885; one is reported to have been captured at Blackpool, about 1860; andone at Wrington, about twelve miles north of Bristol, in 1895 or 1896.
Brown Argus.Egg, natural size and enlarged; caterpillar and chrysalis;(a) Egg of "Scotch Argus" enlarged.
Brown Argus.Egg, natural size and enlarged; caterpillar and chrysalis;(a) Egg of "Scotch Argus" enlarged.
Brown Argus.
Egg, natural size and enlarged; caterpillar and chrysalis;
(a) Egg of "Scotch Argus" enlarged.
Pl.104.Silver-studded Blue.Eggs, natural size and enlarged; caterpillar and chrysalids.
Pl.104.Silver-studded Blue.Eggs, natural size and enlarged; caterpillar and chrysalids.
Pl.104.
Silver-studded Blue.
Eggs, natural size and enlarged; caterpillar and chrysalids.
Larger ImagePl.105.Silver-studded Blue. 1, 2, 3male; 4, 5, 6female.Brown Argus. 10, 12male; 7, 8, 9female; 11, 13, 14male (Durham); 15male, 16, 17female (Scotland).
Pl.105.Silver-studded Blue. 1, 2, 3male; 4, 5, 6female.Brown Argus. 10, 12male; 7, 8, 9female; 11, 13, 14male (Durham); 15male, 16, 17female (Scotland).
Pl.105.
Silver-studded Blue. 1, 2, 3male; 4, 5, 6female.
Brown Argus. 10, 12male; 7, 8, 9female; 11, 13, 14
male (Durham); 15male, 16, 17female (Scotland).
The following details of the early stages are obtained from Mr. Frohawk's life-history of the species published in theEntomologistfor October, 1904. The egg (Plate102,figured from a photomicrograph by Mr. Tonge) is of a pale greenish-blue, but varies both in the extent of the ground colour and in the structure of the reticulations, which are white, resembling frosted glass.
The full-grown caterpillar (August 23rd) measures 3/8 inch in length. It is of the usual wood-louse shape, with only a very shallow furrow on the back, bordered on each side by a fringe of spinous bristles, which vary in length; the whole surface is densely studded with shorter but similarly formed whitish or brownish bristles. The ground colour is pale green, with a darker green stripe along the centre of the back, and fainter green oblique stripes on the sides. The head is black and shining, and is hidden under the first ring when the caterpillar is not feeding or moving about.
The caterpillars hatched on July 30th, from eggs that were laid in the South of France on July 24th, and were reared on bird's-foot trefoil (Lotus corniculatus), of which they ate the flowers, seeds, and leaves.
The chrysalis, which is attached to the food-plant by a silk pad at the tail and a thread round the body, is pale green and very finely reticulated; the wing-cases are rather whiter green, sprinkled with minute black specks, and the veins are white; there is a blackish line along the centre of the back, but this is only well defined on the head and thorax. The whole surface, except the wings, is sprinkled with slightly curved and moderately long white hairs.
The butterfly emerges in about ten to fourteen days, according to temperature.
The male is violet-blue with the veins rather darker; theouter margin is narrowly bordered with blackish, and there are some black dots on the outer margin of the hind wings; the fringes are white, and there is a slender tail on the hind wings. The female is brownish, tinged with violet towards the base; the hind wings have black spots on the outer margin, and some of these are inwardly edged with orange; the tails are slightly longer than those of the male.
All the available information concerning the occurrence of this species in England has already been given. No doubt the localities from which specimens were recorded have been closely investigated during the past twenty years, but no further captures of this butterfly have been recorded. This seems to indicate that it is not really indigenous, but that its presence here may possibly have been due to accidental introduction.
The spring form,polysperchon, is smaller than the specimens occurring in the summer, but so far that form has not been seen in England.
The species is widely distributed over Central and Southern Europe, and its range extends through Northern Asia to Amurland, Corea, and Japan. It is also represented in Northern and Central America by var.comyntas, and has been recorded from Australia.
The male of this butterfly (Plate105) is purplish-blue with a black border on the outer margins, and sometimes black dots on that of the hind wings. The female is sooty-brown, powdered to a greater or lesser extent with blue scales on the basal area; there is generally a series of orange marks forming a more or less complete band on the outer margin of the hind wings, and sometimes on the fore wings also. The under side is bluish-grey in the male, and brownish-grey in the female; the black spots are ringed with white, and on thefore wings there is one at the end of the discal cell and a series of seven beyond; the hind wings have from three to five spots before the discal spot, and a curved series of seven beyond; there is a black-edged orange band on all the wings, and beyond this on the hind wings there is a series of metallic blue centred spots; hence the English name of the butterfly, given to it by Moses Harris, which is certainly more suitable than Petiver's "Lead Argus."
In a general way the male is rather larger than the female, but this is not invariably the case. The colour of the male varies in shade, and very occasionally, perhaps, is of a lilac tint; the border varies in width, and is sometimes reduced to a mere line. In the female the orange marks may be of a brownish or yellowish tint, and now and then there may be a series of wedge-shaped blue spots above these marks on the hind wings. On the under side there is a good deal of modification of the black spots as regards size and shape, and occasionally there is at least one extra spot on the fore wings placed between the discal spot and the base of the wing; white markings sometimes appear on the fore wings between the outer series of black spots and the orange band, and with this there is generally a white band in a similar position on the hind wings. Female specimens with splashes of the male colour on one or more of the wings have been obtained, and, more rarely, examples entirely male on one side and female on the other have been recorded.
Frohawk states that the egg both in colour and texture, resembles white porcelain; "all the depths produce a deep purplish-grey shade. The ova are deposited singly, and adhere firmly to the receptacle."
Caterpillars hatched out from eggs, laid the previous summer, on April 1st to 3rd. They were reared on gorse (Ulex europæus), pupated towards the end of June, and the first butterfly, a male, appeared on July 10th.
The caterpillar figured on Plate104,when full grown, was reddish-brown, finely dotted with white, and from each dot a tiny hair arose; the stripe on the back and line on the side were black edged with white, head black and shining. This caterpillar was found on the last day of May, crawling on the ground under heather at Oxshott. It was then about half-grown, and was reared on heather, pupated in due course, and produced a female butterfly on July 11th.
The chrysalis, of which two figures are given, had a pale brownish and rather shining head; the body was brown with a darker line on the back; the thorax and wing-cases dull yellowish-green, the former rather glossy. It was placed in an angle formed by a side and the floor of the cage, lying quite flat and secured by silken threads, which, owing to position, I was unable to examine. Some of the caterpillars that Mr. Frohawk reared were pale green with a dark purplish stripe on the back. Another food-plant is bird's-foot vetch (Ornithopus perpusillus).
The butterfly is on the wing in July and August, and seems to be more often found on sandy heaths than elsewhere. It is especially common, in some years, in the heather-clad districts of Surrey and Hampshire, as well as other counties in England. In Norfolk and Suffolk it is said to be common, but scarce in Gloucestershire and Somersetshire. Its range extends through the greater part of England and Wales, and into Scotland as far as Perthshire. Specimens from the northwest coast of Wales are said to be larger than those from inland localities.
As regards Ireland, there is only Birchall's record, "The Murrough of Wicklow, and near Rostrevor," in evidence of the butterfly occurring in that country at all.
Abroad, it appears to range pretty well over the whole of Europe, and through Asia eastward to Siberia, Corea, and Japan.
Fore wings blackish or sooty-brown with a black discal spot, and a row of reddish-orange spots on the outer margin of all the wings; the fringes are white, sometimes with blackish interruptions. The under side is greyish or greyish-brown, and the black spots are distinctly ringed with white. On the fore wing there are seven of these spots, one at the end of the cell, and the others in an irregular series beyond; the last in this series is sometimes double, or it may be absent. On the hind wings the spots comprise a series of four preceding the white discal mark, and a series of seven beyond; the second spot in this series is placed directly under the first, forming a colon-like mark, and this character will help to distinguish the Brown Argus from the blackish or brown females of the next species.
The female has larger orange markings, and the outline of the fore wings is rather rounder on the outer margin, otherwise the sexes are very similar.
The orange spots referred to in the male are sometimes absent towards the tips of the fore wings, and in this respect lead up to the form known as the Durham Argus (var.salmacis, Stephens), which is blackish above and ochreous-brown below; the black spots on the under side are much smaller then in typical specimens, and some may be absent altogether. The male has a black discal spot, and the female a white one, on the upper side of the fore wings; the hind wings have a red or orange band on both surfaces. Sometimes the male also has a white spot on the fore wings. Specimens with the orange spots on upper side almost entirely absent are referable to var.allous.
Artaxerxesis the form occurring in Scotland, and is known as the "Scotch White Spot." Both sexes have a conspicuouswhite discal spot on the fore wings, and the spots on the under side are white, and rarely centred with black. In var.quadripuncta, Tutt, all four wings have a white discal spot above. Occasionally an odd specimen with white discal spots is found in the south.
Figures of the butterfly will be found on Plate105,and of its life history on Plate104;the upper egg is that from a typical female, and the lower one was laid by a femaleartaxerxes.
The egg, which is whitish, with a faint greyish tinge, is laid on the upper side of a young leaf of the rock-rose (Helianthemum chamæcistus). The caterpillar has a black shining head; the body is green with whitish hairs, a pinkish line along the back, a whitish one bordered with pinkish along the sides; the green colour becomes dingy as the caterpillar matures. The chrysalis is obscure yellowish-green, the front of the thorax is edged with pinkish, and there are bands of the same colour on the back and sides of the body; the thorax and the wing-cases are rather glossy. Held in position by a few silken threads between leaves of the food-plant.
The ordinary form of the butterfly is on the wing in May and June, and again in August. It is widely distributed throughout the southern half of England, and also in Wales.
Although chiefly associated with rock-rose, especially in chalky districts, it occurs too among stork's-bill (Erodium cicutarium), upon which plant the caterpillar also feeds, in sandy places inland as well as on the coast.
Caterpillars from the first flight of butterflies may be found in July, and those from the second flight hibernate and feed up in April.
The butterfly has a marked liking for roosting on the flower stems of long grasses, and quite a number may often be found resting together towards sundown, or on dull days, in sheltered hollows. Sometimes several specimens of this species and of the Common Blue may be found on the same perch. It israther less frequently seen in the Midland counties, but it is more or less common in some parts of Derbyshire, Yorkshire, and Lancashire.