U. barnesianusLea, ’38.U. meredithiLea, ’58.U. pudicusLea, ’60.U. LyoniLea, ’65.U. tellicoensisLea, ’72.U. lenticularisLea, ’72.
As the normal (most abundant) forms we may regardU. meredithi,pudicusandlenticularis, which differ from each other only in the development of the rays (topotypes examined).U. barnesianusis a slightly more elongated individual, with poorly developed rays.U. tellicoensis(topotypes examined) is alenticularisslightly more swollen; andU. Lyoniforms the transition toward var.tumescens, having a little more elevated beaks, greater obesity, and rather distinct rays.