MONKEY-FLOWER(Minulus ringens) Figwort family
Flowers - Purple, violet, or lilac, rarely whitish; about 1 in. long, solitary, borne on slender footstems from axils of upper leaves. Calyx prismatic, 5-angled, 5-toothed; corolla irregular, tubular, narrow in throat, 2-lipped; upper lip 2-lobed, erect; under lip 3-lobed, spreading; 4 stamens, a long and a short pair, inserted on corolla tube; pistil with 2-lobed, plate-like stigma. Stem: Square, erect, usually branched, 1 to 3 ft. high. Leaves: Opposite, oblong to lance-shaped, saw-edged, mostly seated on stem. Preferred Habitat - Swamps, beside streams and ponds. Flowering Season - June-September. Distribution - Manitoba, Nebraska, and Texas, eastward to Atlantic Ocean.
No wader is the square-stemmed Monkey-flower whose grinning corolla peers at one from grassy tuffets in swamps, from the brookside, the springy soil of low meadows, and damp hollows beside the road; but moisture it must have to fill its nectary and to soften the ground for the easier transit of its creeping rootstock. Imaginative eyes see what appears to them the gaping (ringens) face of a little ape or buffoon (mimulus) in this common flower whose drolleries, such as they are, call forth the only applause desired - the buzz of insects that become pollen-laden during the entertainment.
Now the advanced stigma of this flower is peculiarly irritable, and closes up on contact with an incoming visitor's body, thus exposing the pollen-laden anthers behind it, and, except in rare cases, preventing self-fertilization. Delpino was the first to guess what advantage so sensitive a stigma might mean. Probably the smaller bees find the tube too long for their short tongues. The yellow palate, which partially guards the entrance to the nectary from pilferers, of course serves also as a pathfinder to the long-tongued bees.
AMERICAN BROOKLIME(Veronica Americana) Figwort family
Flowers - Light blue to white, usually striped with deep blue or purple structure of flower similar to that of V. officinalis, but borne in long, loose racemes branching outward on stems that spring from axils of most of the leaves. Stem: Without hairs, usually branched, 6 in. to 3 ft. long, lying partly on ground and rooting from lower joints. Leaves: Oblong, lance-shaped, saw-edged, opposite, petioled, and lacking hairs; 1 to 3 in. long, 1/4 to 1 in. wide. Fruit: A nearly round, compressed, but not flat, capsule with flat seeds in 2 cells. Preferred Habitat - In brooks, ponds, ditches, swamps. Flowering Season - April-September. Distribution - From Atlantic to Pacific, Alaska to California and New Mexico, Quebec to Pennsylvania.
This, the perhaps most beautiful native speedwell, whose sheets of blue along the brookside are so frequently mistaken for masses of forget-me-nots by the hasty observer, of course shows marked differences on closer investigation; its tiny blue flowers are marked with purple pathfinders, and the plant is not hairy, to mention only two. But the poets of England are responsible for most of whatever confusion stills lurks in the popular mind concerning these two flowers. Speedwell, a common medieval benediction from a friend, equivalent to our farewell or adieu, and forget - me-not of similar intent, have been used interchangeably by some writers in connection with parting gifts of small blue flowers. It was the germander speedwell that in literature and botanies alike was most commonly known as the forget-me-not for over two hundred years, or until only fifty years ago. When the "Mayflower" and her sister ships were launched; "Speedwell" was considered a happier name for a vessel than it proved to be.
The WATER SPEEDWELL, or PIMPERNEL (V. Anagallis-aquatica), differs from the preceding chiefly in having most of its leaves seated on the stalk, only the lower ones possessing stems, and those short ones. In autumn the increased growth of sterile shoots from runners produce almost circular leaves, often two inches broad, a certain aid to identification.
Another close relation, the MARSH or SKULLCAP SPEEDWELL (V. scutellata), on the other hand, has long, very slender, acute leaves, their teeth far apart; and as these three species are the only members of their clan likely to be found in watery places within our limits, a close examination of the leaves of any water-loving plant bearing small four-lobed blue flowers, usually marked with lines of a deeper blue or purple, should enable one to correctly name the species. None of these blossoms can be carried far after being picked; they have a tantalizing habit of dropping off, leaving a bouquet of tiny green calices chiefly.
Many kinds of bees, wasps, flies, and butterflies fertilize all these little flowers, which are first staminate, then pistillate, simply by crawling over them in search of nectar.
COMMON SPEEDWELL; FLUELLIN; PAUL'S BETONY; GROUND-HELE(Veronica officinalis) Figwort family
Flowers - Pale blue, very small, crowded on spike-like racemes from axils of leaves, often from alternate axils. Calyx 4-parted; corolla of 4 lobes, lower lobe commonly narrowest ; 2 divergent stamens inserted at base and on either side of upper corolla lobe ; a knob-like stigma on solitary pistil. Stem: From 3 to 10 in. long, hairy, often prostrate, and rooting at joints. Leaves: Opposite, oblong, obtuse, saw-edged, narrowed at base. Fruit: Compressed heart-shaped capsule, containing numerous flat seeds. Preferred Habitat - Dry fields, uplands, open woods. Flowering Season - May-August. Distribution - From Michigan and Tennessee eastward, also from Ontario to Nova Scotia. Probably an immigrant from Europe and Asia.
An ancient tradition of the Roman Church relates that when Jesus was on His way to Calvary, He passed the home of a certain Jewish maiden, who, when she saw the drops of agony on His brow, ran after Him along the road to wipe His face with her kerchief. This linen, the monks declared, ever after bore the impress of the sacred features - vera iconica, the true likeness. When the Church wished to canonize the pitying maiden, an abbreviated form of the Latin words was given her, St. Veronica, and her kerchief became one of the most precious relics at St. Peter's, where it is said to be still preserved. Medieval flower lovers, whose piety seems to have been eclipsed only by their imaginations, named this little flower from a fancied resemblance to the relic. Of course, special healing virtue was attributed to the square of pictured linen, and since all could not go to Rome to be cured by it, naturally the next step was to employ the common, wayside plant that bore the saint's name. Mental healers will not be surprised to learn that because of the strong popular belief in its efficacy to cure all fleshly ills, it actually seemed to possess miraculous powers. For scrofula it was said to be the infallible remedy, and presently we find Linnaeus grouping this flower, and all its relatives under the family name of Scrofulariaceae. "What's in a name?" Religion, theology, medicine, folk-lore, metaphysics, what not?
One of the most common wild flowers in England is this same familiar little blossom of that lovely shade of blue known by Chinese artists as "the sky after rain." "The prettiest of all humble roadside flowers I saw," says Burroughs, in "A Glance at British Wild Flowers." "It is prettier than the violet, and larger and deeper colored than our houstonia. It is a small and delicate edition of our hepatica, done in indigo blue, and wonted to the grass in the fields and by the waysides.
'The little speedwell's darling blue'
sings Tennyson. I saw it blooming with the daisy and buttercup upon the grave of Carlyle. The tender human and poetic element of his stern, rocky nature was well expressed by it."
Only as it grows in masses is the speedwell conspicuous - a sufficient reason for its habit of forming colonies and of gathering its insignificant blossoms together into dense spikes, since by these methods it issues a flaunting advertisement of its nectar. The flower that simplifies dining for insects has its certain reward in rapidly increased and vigorous descendants. To save repetition, the reader interested in the process of fertilization is referred to the account of the Maryland figwort, since many members of the large family to which both belong employ the same method of economizing pollen and insuring fertile seed. In this case visitors have only to crawl over the tiny blossoms.
>From Labrador to Alaska, throughout almost every section of the United States, in South America, Europe, and Asia, roams the THYME-LEAVED SPEEDWELL (V. serpyllifolia), by the help of its numerous flat seeds, that are easily transported on the wind, and by its branching stem, that lies partly on the ground, rooting where the joints touch earth. The small oval leaves, barely half an inch long, grow in pairs. The tiny blue, or sometimes white, flowers, with dark pathfinders to the nectary, are borne on spike-like racemes at the ends of the stem and branches that rear themselves upward in fields and thickets to display their bloom before the passing bee.
PALE, or NAKED, or ONE-FLOWERED BROOM-RAPE (Thalesia uniflora; Aphyllon uniflorum of Gray) Broom-rape family
Flowers - Violet, rarely white, delicately fragrant, solitary at end of erect, glandular peduncles. Calyx hairy, bell-shaped, 5-toothed, not half the length of corolla, which is 1 in. or less long, with curved tube spreading into 2 lips, 5-lobed, yellow-bearded within; 4 stamens, in pairs, inserted on tube of corolla ; 1 pistil. Stem: About 1 in. long, scaly, often entirely underground; the 1 to 4 brownish scape-like peduncles, on which flowers are borne, from 3 to 8 in. high. Leaves: None. Fruit: An elongated, egg-shaped, 1-celled capsule containing numerous seeds. Preferred Habitat - Damp woods and thickets. Flowering Season - April-June. Distribution - British Possessions and United States from coast to coast, southward to Virginia, and Texas.
A curious, beautiful parasite, fastened on the roots of honest plants from which it draws its nourishment. The ancestors of this species, having deserted the path of rectitude ages ago to live by piracy, gradually lost the use of their leaves, upon which virtuous plants depend as upon a part of their digestive apparatus; they grew smaller and smaller, shriveled and dried, until now that the one-flowered broom-rape sucks its food, rendered already digestible through another's assimilation, no leaves remain on its brownish scapes. Disuse of any talent in the vegetable kingdom, as in the spiritual, leads to inevitable loss: "Unto every one which hath shall be given; and from him that hath not, even that he hath shall be taken away."
HAIRY RUELLIA(Ruellia ciliosa) Acanthus family
Flowers - Pale violet blue, showy, about 2 in. long, solitary or clustered in the axils or at the end of stem. Calyx of 5 bristle-shaped hairy segments; corolla with very slender tube expanding above in 5 nearly equal obtuse lobes; stamens 4; pistil with recurved style. Stem: Hairy, especially above, erect, 1 to 2 1/2 ft. high. Leaves: Opposite, oblong, narrowed at apex, entire, covered with soft white hairs. Preferred Habitat - Dry soil. Flowering Season - June-September. Distribution - New Jersey southward to the Gulf and westward to Michigan and Nebraska.
Many charming ruellias from the tropics adorn hothouses and window gardens in winter; but so far north as the New Jersey pine barrens, and westward where killing frosts occur, this perennial proves to be perfectly hardy. In addition to its showy blossoms, which so successfully invite insects to transfer their pollen, thereby counteracting the bad effects of close inbreeding, the plant bears inconspicuous cleistogamous or blind ones also. These look like arrested buds that never open; but, being fertilized with their own pollen, ripen abundant seed nevertheless.
One frequently finds holes bitten in these flowers, as in so many others long of tube or spur. Bumblebees, among the most intelligent and mischievous of insects, are apt to be the chief offenders; but wasps are guilty too, and the female carpenter bee, which ordinarily slits holes to extract nectar, has been detected in the act of removing circular pieces of the corolla from this ruellia with which to plug up a thimble-shaped tube in some decayed tree. Here she deposits an egg on top of a layer of baby food, consisting of a paste of pollen and nectar, and seals up the nursery with another bit of leaf or flower, repeating the process until the long tunnel is filled with eggs and food for larvae. Then she dies, leaving her entire race apparently extinct, and living only in embryo for months. This is the bee which commonly cuts her round plugs from rose leaves.
The SMOOTH RUELLIA (R. strepens), an earlier bloomer than the preceding, and with a more southerly range, has a shorter, thicker tube to its handsome blue flower, and lacks the hairs which guard its relative from crawling pilferers.
BLUETS; INNOCENCE; HOUSTONIA; QUAKER LADIES; QUAKER BONNETS;VENUS' PRIDE(Houstonia caerulea) Madder family
Flowers - Very small, light to purplish blue or white, with yellow center, and borne at end of each erect slender stem that rises from 3 to 7 in. high. Corolla funnel-shaped, with 4 oval, pointed, spreading lobes that equal the slender tube in length; rarely the corolla has more divisions; 4 stamens inserted on tube of corolla; 2 stigmas; calyx 4-lobed. Leaves: Opposite, seated on stem, oblong, tiny; the lower ones spatulate. Fruit: A 2-lobed pod, broader than long, its upper half free from calyx; seeds deeply concave. Root stock: Slender, spreading, forming dense tufts. Preferred Habitat - Moist meadows, wet rocks and banks. Flowering Season - April-July, or sparsely through summer. Distribution - Eastern Canada and United States west to Michigan, south to Georgia and Alabama.
Millions of these dainty wee flowers, scattered through the grass of moist meadows and by the wayside, reflect the blue and the serenity of heaven in their pure, upturned faces. Where the white variety grows, one might think a light snowfall had powdered the grass, or a milky way of tiny floral stars had streaked a terrestrial path. Linnaeus named the flower for Dr. Houston, a young English physician, botanist, and collector, who died in South America in 1733, after an exhausting tramp about the Gulf of Mexico.
To secure cross-fertilization, the object toward which so much marvelous floral organism is directed, this little plant puts forth two forms of blossoms - one with the stamens in the lower portion of the corolla tube, and the stigmas exserted; the other form with the stigmas below, and the stamens elevated to the mouth of the corolla. But the two kinds do not grow in the same patch, seed from either producing after its kind. Many insects visit these blossoms, but chiefly small bees and butterflies. Conspicuous among the latter is the common little meadow fritillary (Brenthis bellona), whose tawny, dark-speckled wings expand and close in apparent ecstasy as he tastes the tiny drop of nectar in each dainty enameled cup. Coming to feast with his tongue dusted from anthers nearest the nectary, he pollenizes the large stigmas of a short-styled blossom without touching its tall anthers. But it is evident that he could not be depended on to fertilize the long-styled form, with its smaller stigma, because of this ability to insert his slender tongue from the side where it avoids contact. Flies and beetles enter the blossoms, but small bees are best adapted as all-round benefactors. This simple-looking blossom, that measures barely half an inch across, is clever enough to multiply its lovely species a thousand fold, while many a larger, and therefore one might suppose a wiser, flower dwindles toward extinction.
John Burroughs found a single bluet in blossom one January, near Washington, when the clump of earth on which it grew was frozen solid. A pot of roots gathered in autumn and placed in a sunny window has sent up a little colony of star-like flowers throughout a winter.
WILD, COMMON, or CARD TEASEL; GYPSY COMBS(Dipsacus sylvestris) Teasel family
Flowers - Purple or lilac, small, packed in dense, cylindric heads, 3 to 4 in. long; growing singly on ends of footstalks, the flowers set among stiffly pointed, slender scales. Calyx cup-shaped, 4-toothed. Corolla 4-lobed; stamens 4; leaves of involucre, slender, bristled, curved upward as high as flower-head or beyond. Stems: 3 to 6 ft. high, stout, branched, leafy, with numerous short prickles. Leaves: Opposite, lance-shaped, seated on stem, with bristles along the stout midrib. Preferred Habitat - Roadsides and waste places. Flowering Season - July-September. Distribution - Maine to Virginia, westward to Ontario and the Mississippi. Europe and Asia.
Manufacturers find that no invention can equal the natural teasel head for raising a nap on woolen cloth, because it breaks at any serious obstruction, whereas a metal substitute, in such a case, tears the material. Accordingly, the plant is largely cultivated in the west of England, and quantities that have been imported from France and Germany may be seen in wagons on the way to the factories in any of the woolen-trade towns. After the flower-heads wither, the stems are cut about eight inches long, stripped of prickles, to provide a handle, and after drying, the natural tool is ready for use.
Bristling with armor, the teasel is not often attacked by browsing cattle. Occasionally even the upper leaf surfaces are dotted over with prickles enough to tear a tender tongue. This is a curious feature, for prickles usually grow out of veins. In the receptacle formed where the bases of the upper leaves grow together, rain and dew are found collected - a certain cure for warts, country people say. Venus' Cup, Bath, or Basin, and Water Thistle, are a few of the teasel's folk names earned by its curious little tank. In it many small insects are drowned, and these are supposed to contribute nourishment to the plant; for Mr. Francis Darwin has noted that protoplasmic filaments reach out into the liquid.
Owing to the stiff spines which radiate from the flower cluster, the bumblebees, which principally fertilize it, can reach the florets only with their heads, and not pollenize them by merely crawling over them as in the true compositae. But by first maturing its anthers, then when they have shed their pollen, elevating its stigmas, the teasel prevents self-fertilization.
HAREBELL or HAIRBELL; BLUE BELLS of SCOTLAND; LADY'S THIMBLE(Campanula rotundifolia) Bellflower family
Flowers - Bright blue or violet blue, bell-shaped, 1/2 in. long or over, drooping from hair-like stalks. Calyx of 5-pointed, narrow, spreading lobes; slender stamens alternate with lobes of corolla, and borne on summit of calyx tube, which is adherent to ovary; pistil with 3 stigmas in maturity only. Stem: Very slender, 6 in. to 3 ft. high, often several from same root; simple or branching. Leaves: Lower ones nearly round, usually withered and gone by flowering season; stem leaves narrow, pointed, seated on stem. Fruit: An egg-shaped, pendent, 3-celled capsule with short openings near base; seeds very numerous, tiny. Preferred Habitat - Moist rocks, uplands. Flowering Season - June-September. Distribution - Arctic regions of Europe, Asia, and America; southward on this continent, through Canada to New Jersey and Pennsylvania; westward to Nebraska, to Arizona in the Rockies, and to California in the Sierra Nevadas.
The inaccessible crevice of a precipice, moist rocks sprayed with the dashing waters of a lake or some tumbling mountain stream, wind-swept upland meadows, and shady places by the roadside may hold bright bunches of these hardy bells, swaying with exquisite grace on tremulous, hair-like stems that are fitted to withstand the fiercest mountain blasts, however frail they appear. How dainty, slender, tempting these little flowers are! One gladly risks a watery grave or broken bones to bring down a bunch from its aerial cranny.
It was a long stride forward in the evolutionary scale when the harebell welded its five once separate petals together; first at the base, then farther and farther up the sides, until a solid bell-shaped structure resulted. This arrangement which makes insect fertilization a more certain process because none of the pollen is lost through apertures, and because the visitor must enter the flower only at the vital point where the stigmas come in contact with his pollen-laden body, has given to all the flowers that have attained to it, marked ascendency.
Like most inverted blossoms, the harebell hangs its head to protect its nectar and pollen, not only from rain, but from the intrusion of undesirable crawling insects which would simply brush off its pollen in the grass before reaching the pistil of another flower, and so defeat cross-fertilization, the end and aim of so many blossoms. Advertising for winged insects by its bright color, the harebell attracts bees, butterflies, and many others. These visitors cannot well walk on the upright petals, and sooner or later must clasp the pistil if they would secure the nectar secreted at the base. In doing so, they will dust themselves and the immature pistil with the pollen from the surrounding anthers; but a newly opened flower is incapable of fertilization. The pollen, although partially discharged in the unopened bud, is prevented from falling out by a coat of hairs on the upper part of the style. By the time all the pollen has been removed by visitors, however, and the stamens which matured early have withered, the pistil has grown longer, until it looks like the clapper in a bell; the stigma at its top has separated into three horizontal lobes which, being sticky on the under side, a pollen-laden insect on entering the bell must certainly brush against them and render them fertile. But bumblebees, its chief benefactors, and others may not have done their duty by the flower; what then? Why, the stigmas in that case finally bend backward to reach the left over pollen, and fertilize themselves, obviously the next best thing for them to do. How one's reverence increases when one begins to understand, be it ever so little of, the divine plan!
"Probably the most striking blue and purple wild flowers we have," says John Burroughs, "are of European origin. These colors, except with the fall asters and gentians, seem rather unstable in our flora." This theory is certainly borne out in the case of the RAMPION, EUROPEAN, or CREEPING BELLFLOWER (C. rapunculoides), now detected in the act of escaping from gardens from New Brunswick to Ontario, Southern New York, Pennsylvania, and Ohio, and making itself very much at home in our fields and along the waysides. Compared with the delicate little harebell, it is a plant of rank, rigid habit. Its erect, rather stout stem, set with elongated oval, hairy, alternate leaves, and crowned with a one-sided raceme of widely expanded, purple-blue bells rising about two feet above the ground, has little of the exquisite grace of its cousin. It blooms from July to September. This is the species whose roots are eaten by the omnivorous European peasant.
One of the few native campanulas, the TALL BELLFLOWER (C. Americana), waves long, slender wands studded with blue or sometimes whitish flowers high above the ground of moist thickets and woods throughout the eastern half of this country, but rarely near the sea. Doubtless the salt air, which intensifies the color of so many flowers, would brighten its rather slatey blue. The corolla, which is flat, round, about an inch across, and deeply cleft into five pointed petals, has the effect of a miniature pinwheel in motion. Mature flowers have the style elongated, bent downward, then curved upward, that the stigmas may certainly be in the way of the visiting insect pollen-laden from an earlier bloomer, and be cross-fertilized. The larger bees, its benefactors, which visit it for nectar, touch only the upper side of the style, on which they must alight; but the anthers waste pollen by shedding it on all sides. No insect can take shelter from rain or pass the night in this flower, as he frequently does in its more hospitable relative, the harebell. English gardeners, more appreciative than our own of our native flora, frequently utilize this charming plant in their rockwork, increasing their stock by a division of the dense, leafy rosettes.
VENUS' LOOKING-GLASS; CLASPING BELLFLOWER(Legouzia perfoliata; Specularia perfoliata of Gray)Bellflower family
Flowers - Violet blue, from 1/2 to 3/4 in. across; solitary or 2 or 3 together, seated, in axils of upper leaves. Calyx lobes varying from 3 to 5 in earlier and later flowers, acute, rigid; corolla a 5-spoked wheel; 5 stamens; pistil with 3 stigmas. Stem: 6 in. to 2 ft. long, hairy, densely leafy, slender, weak. Leaves: Round, clasped about stem by heart-shaped base. Preferred Habitat - Sterile waste places, dry woods. Flowering Season - May-September. Distribution - From British Columbia, Oregon, and Mexico, east to Atlantic Ocean.
At the top of a gradually lengthened and apparently overburdened leafy stalk, weakly leaning upon surrounding vegetation, a few perfect blossoms spread their violet wheels, while below them insignificant earlier flowers, which, although they have never opened, nor reared their heads above the hollows of the little shell-like leaves where they lie secluded, have, nevertheless, been producing seed without imported pollen while their showy sisters slept. But the later blooms, by attracting insects, set cross-fertilized seed to counteract any evil tendencies that might weaken the species if it depended upon self-fertilization only. When the European Venus' looking-glass used to be cultivated in gardens here, our grandmothers tell us it was altogether too prolific, crowding out of existence its less fruitful, but more lovely, neighbors.
The SMALL VENUS' LOOKING-GLASS (L. biflora), of similar habit to the preceding, but with egg-shaped or oblong leaves seated on, not clasping, its smooth and very slender stem, grows in the South and westward to California.
GREAT LOBELIA; BLUE CARDINAL-FLOWER(Lobelia syphilitica) Bellflower family
Flowers - Bright blue, touched with white, fading to pale blue, about 1 in. long, borne on tall, erect, leafy spike. Calyx 5-parted, the lobes sharply cut, hairy. Corolla tubular, open to base on one side, 2-lipped, irregularly 5-lobed, the petals pronounced at maturity only. Stamens 5, united by their hairy anthers into a tube around the style; larger anthers smooth. Stem: 1 to 3 ft. high, stout, simple, leafy, slightly hairy. Leaves: Alternate, oblong, tapering, pointed, irregularly toothed, 2 to 6 in. long, 1/2 to 2 in. wide. Preferred Habitat - Moist or wet soil; beside streams. Flowering Season - July-October Distribution - Ontario and northern United States west to Dakota, south to Kansas and Georgia.
To the evolutionist, ever on the lookout for connecting links, the lobelias form an interesting group, because their corolla, slit down the upper side and somewhat flattened, shows the beginning of the tendency toward the strap or ray flowers that are nearly confined to the composites of much later development, of course, than tubular single blossoms. Next to massing their flowers in showy heads, as the composites do, the lobelias have the almost equally advantageous plan of crowding theirs along a stem so as to make a conspicuous advertisement to attract the passing bee and to offer him the special inducement of numerous feeding places close together.
The handsome GREAT LOBELIA, constantly and invidiously compared with its gorgeous sister the cardinal flower, suffers unfairly. When asked what his favorite color was, Eugene Field replied: "Why, I like any color at all so long as it's red!" Most men, at least, agree with him, and certainly hummingbirds do; our scarcity of red flowers being due, we must believe, to the scarcity of hummingbirds, which chiefly fertilize them. But how bees love the blue blossoms!
There are many cases where the pistil of a flower necessarily comes in contact with its own pollen, yet fertilization does not take place, however improbable this may appear. Most orchids, for example, are not susceptible to their own pollen. It would seem as if our lobelia, in elevating its stigma through the ring formed by the united anthers, must come in contact with some of the pollen they have previously discharged from their tips, not only on the bumblebee that shakes it out of them when he jars the flower, but also within the tube. But when the anthers are mature, the two lobes of the still immature stigma are pressed together, and cannot be fertilized. Nevertheless, the hairy tips of some of the anthers brush off the pollen grains that may have lodged on the stigma as it passes through the ring in its ascent, thus making surety doubly sure. Only after the stigma projects beyond the ring of anthers does it expand its lobes, which are now ready to receive pollen brought from another later flower by the incoming bumblebee to which it is adapted.
Linnaeus named this group of plants for Matthias de l'Obel, a Flemish botanist, or herbalist more likely, who became physician to James I. of England.
Preferably in dry, sandy soil or in meadows, and over a wide range, the slender, straight shoots of PALE SPIKED LOBELIA (L. spicata) bloom early and throughout the summer months, the inflorescence itself sometimes reaching a height of two feet. At the base of the plant there is usually a tuft of broadly oblong leaves; those higher up narrow first into spoon-shaped, then into pointed, bracts, along the thick and gradually lengthened spike of scattered bloom. The flowers are oft en pale enough to be called white. Like their relatives, they first ripen their anthers to prevent self-fertilization.
The lithe, graceful little BROOK LOBELIA (L. Kalmii), whose light-blue flowers, at the end of thread-like footstems, form a loose raceme, sways with a company of its fellows among the grass on wet banks, beside meadow runnels and brooks, particularly in limestone soil, from Nova Scotia to the Northwest Territory and southward to New Jersey. It bears an insignificant capsule, not inflated like the Indian tobacco's; and long, narrow, spoon-shaped leaves. Twenty inches is the greatest height this little plant may hope to attain.
Not only beside water, and in it, but often totally immersed, grows the WATER LOBELIA or GLADIOLE (L. Dortmanna). The slender, hollow, smooth stem rises from a submerged tuft of round, hollow, fleshy leaves longitudinally divided by a partition, and bears at the top a scattered array of pale-blue flowers from August to September.
INDIAN or WILD TOBACCO; GAG-ROOT; ASTHMA-WEED; BLADDER-PODLOBELIA(Lobelia inflata) Bellflower family
Flowers - Pale blue or violet, small, borne at short intervals in spike-like leafy racemes. Calyx 5-parted, its awl-shaped lobes 1/4 in. long, or as long as the tubular, 2-lipped, 5-cleft, corolla that opens to base of tube on upper side. Stamens, 5 united by their hairy anthers into a ring around the 2-lobed style. Stem: From 1 to 3 feet high, hairy, very acrid, much branched, leafy. Leaves: Alternate, oblong or ovate, toothed, the upper ones acute, seated on stem; lower ones obtuse, petioled, to 2 1/2 in. long. Fruit: A much inflated, rounded, ribbed, many seeded capsule. Preferred Habitat - Dry fields and thickets; poor soil. Flowering Season - July-November. Distribution - Labrador westward to the Missouri River, south to Arkansas and Georgia.
The most stupid of the lower animals knows enough to let this poisonous, acrid plant alone; but not so man, who formerly made a quack medicine from it in the days when a drug that set one's internal organism on fire was supposed to be especially beneficial. One taste of the plant gives a realizing sense of its value as an emetic. How the red man enjoyed smoking and chewing the bitter leaves, except for the drowsiness that followed, is a mystery.
On account of the smallness of its flowers and their scantiness, the Indian tobacco is perhaps the least attractive of the lobelias, none of which has so inflated a seed vessel, the distinguishing characteristic of this common plant.
CHICORY; SUCCORY; BLUE SAILORS; BUNK(Cichorium Intybus) Chicory family
Flower-head - Bright, deep azure to gray blue, rarely pinkish or white, 1 to 1 1/2 in. broad, set close to stem, often in small clusters for nearly the entire length; each head a composite of ray flowers only, 5-toothed at upper edge, and set in a flat green receptacle. Stem: Rigid, branching, to 3 ft. high. Leaves: Lower ones spreading on ground, 3 to 6 in. long, spatulate, with deeply cut or irregular edges, narrowed into petioles, from a deep tap-root; upper leaves of stem and branches minute, bract-like. Preferred Habitat - Roadsides, waste places, fields. Flowering Season - July-October. Distribuition - Common in Eastern United States and Canada, south to the Carolinas; also sparingly westward to Nebraska.
At least the dried and ground root of this European invader is known to hosts of people who buy it undisguised or not, according as they count it an improvement to their coffee or a disagreeable adulterant. So great is the demand for chicory that, notwithstanding its cheapness, it is often in its turn adulterated with roasted wheat, rye, acorns, and carrots. Forced and blanched in a warm, dark place, the bitter leaves find a ready market as a salad known as "barbe de Capucin" by the fanciful French. Endive and dandelion, the chicory's relatives, appear on the table too, in spring, where people have learned the possibilities of salads, as they certainly have in Europe.
>From the depth to which the tap-root penetrates, it is not unlikely the succory derived its name from the Latin succurrere = to run under. The Arabic name chicourey testifies to the almost universal influence of Arabian physicians and writers in Europe after the Conquest. As chicoree, achicoria, chicoria, cicorea, chicorie, cichorei, cikorie, tsikorei, and cicorie the plant is known respectively to the French, Spanish, Portuguese, Italians, Germans, Dutch, Swedes, Russians, and Danes.
On cloudy days or in the morning only throughout midsummer the "peasant posy" opens its "dear blue eyes"
"Where tired feetToil to and fro;Where flaunting SinMay see thy heavenly hue,Or weary Sorrow look from theeToward a tenderer blue!"- Margaret Deland.
In his "Humble Bee" Emerson, too, sees only beauty in the "Succory to match the sky;" but, mirabile dictu, Vergil, rarely caught in a prosaic, practical mood, wrote, "And spreading succ'ry chokes the rising field."
IRON-WEED; FLAT TOP(Vernonia Noveboracensis) Thistle family.
Flower-head - Composite of tubular florets only, intense reddish-purple thistle-like heads, borne on short, branched peduncles and forming broad, flat clusters; bracts of involucre, brownish purple, tipped with awl-shaped bristles. Stem: 3 to 9 ft. high, rough or hairy, branched. Leaves: Alternate, narrowly oblong or lanceolate, saw-edged, 3 to 10 in. long, rough. Preferred Habitat - Moist soil, meadows, fields. Flowering Season - July-September. Distribution - Massachusetts to Georgia, and westward to the Mississippi.
Emerson says a weed is a plant whose virtues we have not yet discovered; but surely it is no small virtue in the iron-weed to brighten the roadsides and low meadows throughout the summer with bright clusters of bloom. When it is on the wane, the asters, for which it is sometimes mistaken, begin to appear, but an instant's comparison shows the difference between the two flowers. After noting the yellow disk in the center of an aster, it is not likely the iron-weed's thistle-like head of ray florets only will ever again be confused with it. Another rank-growing neighbor with which it has been confounded by the novice is the Joe Pye weed, a far paler, pinkish flower.
To each tiny floret, secreting nectar in its tube, many insects, attracted by the bright color of the iron-weed standing high above surrounding vegetation, come to feast. Long-lipped bees and flies rest awhile for refreshment, but butterflies of many beautiful kinds are by far the most abundant visitors. Pollen carried out by the long, hairy styles as they extend to maturity must attach itself to their tongues. The tiger swallow-tail butterfly appears to have a special preference for this flower. (See Self-Heal.)
COMMON or SCALY BLAZING STAR; COLIC-ROOT; RATTLESNAKE MASTER;BUTTON SNAKEROOT(Lacinaria squarrosa; Liatris squarrosa of Gray) Thistlefamily
Flower-heads - Composite, about 1 in. long, bright purple or rose purple, of tubular florets only, from an involucre of overlapping, rigid, pointed bracts; each of the few flower-heads from the leaf axils along a slender stem in a wand-like raceme. Stem: 1/2 to 2 ft. high. Leaves: Alternate, narrow, entire. Preferred habitat - Dry, rich soil. Flowering Season - June-September. Distribution - Ontario to the Gulf of Mexico, westward to Nebraska.
Beginning at the top, the apparently fringed flower-heads open downward along the wand, whose length depends upon the richness of the soil. All of the flowers are perfect and attract long-tongued bees and flies (especially Exoprosopa fasciata) and butterflies, which, as they sip from the corolla tube, receive the pollen carried out and exposed on the long divisions of the style. Some people have pretended to cure rattlesnake bites with applications of the globular tuber of this and the next species.
The LARGE BUTTON SNAKEROOT, BLUE BLAZING STAR, or GAY FEATHER (L. scariosa), may attain six feet, but usually not more than half that height; and its round flower-heads normally stand well away from the stout stem on foot-stems of their own. The bristling scales of the involucre, often tinged with purple at the tips, are a conspicuous feature. With much the same range and choice of habitat as the last species, this Blazing Star is a later bloomer, coming into flower in August, and helping the goldenrods and asters brighten the landscape throughout the early autumn. The name of gay feather, miscellaneously applied to several blazing stars, is especially deserved by this showy beauty of the family.
Unlike others of its class, the DENSE BUTTON SNAKEROOT, DEVIL'S BIT, ROUGH or BACKACHE ROOT, PRAIRIE PINE or THROATWORT (L. spicata), the commonest species we have, chooses moist soil, even salt marshes near the coast, and low meadows throughout a range nearly corresponding with that of the scaly blazing star. Resembling its relatives in general manner of growth, we note that its oblong involucre, rounded at the base, has blunt, not sharply pointed, bracts; that the flower-heads are densely set close to the wand for from four to fifteen inches; that the five to thirteen bright rose-purple florets which compose each head occasionally come white; that its leaves are long and very narrow, and that October is not too late to find the plant in bloom.
BLUE and PURPLE ASTERS or STARWORTSThistle family
Evolution teaches us that thistles, daisies, sunflowers, asters, and all the triumphant horde of composites were once very different flowers from what we see today. Through ages of natural selection of the fittest among their ancestral types, having finally arrived at the most successful adaptation of their various parts to their surroundings in the whole floral kingdom, they are now overrunning the earth. Doubtless the aster's remote ancestors were simple green leaves around the vital organs, and depended upon the wind, as the grasses do - a most extravagant method - to transfer their pollen. Then some rudimentary flower changed its outer row of stamens into petals, which gradually took on color to attract insects and insure a more economical method of transfer. Gardeners today take advantage of a blossom's natural tendency to change stamens into petals when they wish to produce double flowers. As flowers and insects developed side by side, and there came to be a better and better understanding between them of each other's requirements, mutual adaptation followed. The flower that offered the best advertisement, as the composites do, by its showy rays; that secreted nectar in tubular flowers where no useless insect could pilfer it; that fastened its stamens to the inside wall of the tube where they must dust with pollen the underside of every insect, unwittingly cross-fertilizing the blossom as he crawled over it; that massed a great number of these tubular florets together where insects might readily discover them and feast with the least possible loss of time - this flower became the winner in life's race. Small wonder that our June fields are white with daisies and the autumn landscape is glorified with goldenrod and asters!
Since North America boasts the greater part of the two hundred and fifty asters named by scientists, and as variations in many of our common species frequently occur, the tyro need expect no easy task in identifying every one he meets afield. However, the following are possible acquaintances to everyone:
In dry, shady places the LARGE or BROAD-LEAVED ASTER (A. macrophyllus), so called from its three or four conspicuous, heart-shaped leaves on long petioles, in a clump next the ground, may be more easily identified by these than by the pale lavender or violet flower-heads of about sixteen rays each which crown its reddish angular stem in August and September. The disk turns reddish brown.
In prairie soil, especially about the edges of woods in western New York, southward and westward to Texas and Minnesota, the beautiful SKY-BLUE ASTER (A. azureus) blooms from August till after frost. Its slender, stiff, rough stem branches above to display the numerous bright blue flowers, whose ten to twenty rays measure only about a quarter of an inch in length. The upper leaves are reduced to small flat bracts; the next are linear; and the lower ones, which approach a heart shape, are rough on both sides, and may be five or six inches long.
Much more branched and bushy is the COMMON BLUE, BRANCHING, WOOD, or HEART-LEAVED ASTER (A. cordifolius), whose generous masses of small, pale lavender flower-heads look like a mist hanging from one to five feet above the earth in and about the woods and shady roadsides from September even to December in favored places.
The WAVY or VARIOUS-LEAVED ASTER or SMALL FLEABANE (A. undulatus) has a stiff, rough, hairy, widely branching stalk, whose thick, rough lowest leaves are heart-shaped and set on long foot-stems; above these, the leaves have shorter stems, dilating where they clasp the stalk; the upper leaves, lacking stems, are seated on it, while those of the branches are shaped like tiny awls. The flowers, which measure less than an inch across, often grow along one side of an axis as well as in the usual raceme. Eight to fifteen pale blue to violet rays surround the disks which, yellow at first, become reddish brown in maturity. We find the plant in dry soil, blooming in September and October.
By no means tardy, the LATE PURPLE ASTER, so-called, or PURPLE DAISY (A. patens), begins to display its purplish-blue, daisy-like flower-heads early in August, and farther north may be found in dry, exposed places only until October. Rarely the solitary flowers, that are an inch across or more, are a deep, rich violet. The twenty to thirty rays which surround the disk, curling inward to dry, expose the vase-shaped, green, shingled cups that terminate each little branch. The thick, somewhat rigid, oblong leaves, tapering at the tip, broaden at the base to clasp the rough, slender stalk. Range similar to the next species.
Certainly from Massachusetts, northern New York, and Minnesota southward to the Gulf of Mexico one may expect to find the NEW ENGLAND ASTER or STARWORT (A. Novae-Angliae) one of the most striking and widely distributed of the tribe, in spite of its local name. It is not unknown in Canada. The branching clusters of violet or magenta-purple flower-heads, from one to two inches across - composites containing as many as forty to fifty purple ray florets around a multitude of perfect five-lobed, tubular, yellow disk florets in a sticky cup - shine out with royal splendor above the swamps, moist fields, and roadsides from August to October. The stout, bristle-hairy stem bears a quantity of alternate lance-shaped leaves lobed at the base where they clasp it.
In even wetter ground we find the RED-STALKED, PURPLE-STEMMED, or EARLY PURPLE ASTER, COCASH, SWANWEED, or MEADOW SCABISH (A. puniceus) blooming as early as July or as late as November. Its stout, rigid stem, bristling with rigid hairs, may reach a height of eight feet to display the branching clusters of pale violet or lavender flowers. The long, blade-like leaves, usually very rough above and hairy along the midrib beneath, are seated on the stem. The lovely SMOOTH or BLUE ASTER (A. laevis), whose sky-blue or violet flower-heads, about one inch broad, are common through September and October in dry soil and open woods, has strongly clasping, oblong, tapering leaves, rough margined, but rarely with a saw-tooth, toward the top of the stem, while those low down on it gradually narrow into clasping wings.
In dry, sandy soil, mostly near the coast, from Massachusetts to Delaware, grows one of the loveliest of all this beautiful clan, the LOW, SHOWY, or SEASIDE PURPLE ASTER (A. spectabilis). The stiff, usually unbranched stem does its best in attaining a height of two feet. Above, the leaves are blade-like or narrowly oblong, seated on the stem, whereas the tapering, oval basal leaves are furnished with long footstems, as is customary with most asters. The handsome, bright, violet-purple flower-heads, measuring about an inch and a half across, have from fifteen to thirty rays, or only about half as many as the familiar New England aster. Season August to November.
The low-growing BOG ASTER (A. nemoralis), not to be confused with the much taller Red-stalked species often found growing in the same swamp, and having, like it, flower-heads measuring about an inch and a half across, has rays that vary from light violet purple to rose pink. Its oblong to lance-shaped leaves, only two inches long at best, taper to a point at both ends, and are seated on the stem. We look for this aster in sandy bogs from New Jersey northward and westward during August and September.
The STIFF or SAVORY-LEAVED ASTER, SANDPAPER, or PINE STARWORT (Ionactis linariifolius), now separated from the other asters into a genus by itself, is a low, branching little plant with no basal leaves, but some that are very narrow and blade-like, rigid, entire and one-nerved, ascending the stiff stems. The leaves along the branches are minute and awl-shaped, like those on a branch of pine. Only from ten to fifteen violet ray flowers (pistillate) surround the perfect disk florets. From Quebec to the Gulf of Mexico, and westward beyond the Mississippi this prim little shrub grows in tufts on dry or rocky soil, and blooms from July to October.
ROBIN'S, or POOR ROBIN'S, or ROBERT'S PLANTAIN; BLUE SPRINGDAISY; DAISY-LEAVED FLEABANE(Erigeron pulchellus; E. bellifolium of Gray) Thistle family
Flower-heads - Composite, daisy-like, 1 to 1/2 in. across; the outer circle of about 50 pale bluish-violet ray florets; the disk florets greenish yellow. Stem: Simple, erect, hairy, juicy, flexible, from 10 in. to 2 ft. high, producing runners and offsets from base. Leaves: Spatulate, in a flat tuft about the root; stem leaves narrow, more acute, seated, or partly clasping. Preferred Habitat - Moist ground, hills, banks, grassy fields. Flowering Season - April-June. Distribution - United States and Canada, east of the Mississippi.
Like an aster blooming long before its season, Robin's plantain wears a finely cut lavender fringe around a yellow disk of minute florets; but one of the first, not the last, in the long procession of composites has appeared when we see gay companies of these flowers nodding their heads above the grass in the spring breezes as if they were village gossips.
Doubtless it was the necessity for attracting insects which led the Robin's plantain and other composites to group a quantity of minute florets, each one of which was once an independent, detached blossom, into a common head. In union there is strength. Each floret still contains, however, its own tiny drop of nectar, its own stamens, its own pistil connected with embryonic seed below; therefore, when an insect alights where he can get the greatest amount of nectar for the least effort, and turns round and round to exhaust each nectary, he is sure to dust the pistils with pollen, and so fertilize an entire flower-head in a trice. The lavender fringe and the hairy involucre and stem serve the end of discouraging crawling insects, which cannot transfer pollen from plant to plant, from pilfering sweets that cannot be properly paid for. Small wonder that, although the composites have attained to their socialistic practices at a comparatively recent day as evolutionists count time, they have become as individuals and as species the most numerous in the world; the thistle family, dominant everywhere, containing not less than ten thousand members.
COMMON or PHILADELPHIA FLEABANE, or SKEVISH (E. Philadelphicus), a smaller edition of Robin's plantain, with a more findely cut fringe, its reddish-purple ray florets often numbering one hundred and fifty, may be found in low fields and woods throughout North America, except in the circumpolar regions.
THISTLES(Carduus) Thistle family
Is land fulfilling the primal curse because it brings forth thistles? So thinks the farmer, no doubt, but not the goldfinches which daintily feed among the fluffy seeds, nor the bees, nor the "painted lady," which may be seen in all parts of the world where thistles grow, hovering about the beautiful rose-purple flowers. In the prickly cradle of leaves, the caterpillar of this thistle butterfly weaves a web around its main food store.
When the Danes invaded Scotland, they stole a silent night march upon the Scottish camp by marching barefoot; but a Dane inadvertently stepped on a thistle, and his sudden, sharp cry, arousing the sleeping Scots, saved them and their country: hence the Scotch emblem.
>From July to November blooms the COMMON, BURR, SPEAR, PLUME, BANK, HORSE, BULL, BLUE, BUTTON, BELL, or ROADSIDE THISTLE (C. lanceolatus or Circium lanceolatum of Gray), a native of Europe and Asia, now a most thoroughly naturalized American from Newfoundland to Georgia, westward to Nebraska. Its violet flower-heads, about an inch and a half across, and as high as wide, are mostly solitary at the ends of formidable branches, up which few crawling creatures venture. But in the deep tube of each floret there is nectar secreted for the flying visitor who can properly transfer pollen from flower to flower. Such a one suffers no inconvenience from the prickles, but, on the contrary, finds a larger feast saved for him because of them. Dense, matted, wool-like hairs, that cover the bristling stems of most thistles, make climbing mighty unpleasant for ants, which ever delight in pilfering sweets. Perhaps one has the temerity to start upward.
"Fain would I climb, yet fear to fall.""If thy heart fail thee, climb not at all,"
might be the ant's passionate outburst to the thistle, and the thistle's reply, instead of a Sir Walter and Queen Elizabeth couplet. Long, lance-shaped, deeply cleft, sharply pointed, and prickly dark green leaves make the ascent almost unendurable; nevertheless the ant bravely mounts to where the bristle-pointed, overlapping scales of the deep green cup hold the luscious flowers. Now his feet becoming entangled in the cottony fibers wound about the scaly armor, and a bristling bodyguard thrusting spears at him in his struggles to escape, death happily releases him. All this tragedy to insure the thistle's cross-fertilized seed that, seated on the autumn winds, shall be blown far and wide in quest of happy conditions for the offspring!
Sometimes the PASTURE or FRAGRANT THISTLE (C. odoratus or C. pumilum of Gray) still further protects its beautiful, odorous purple or whitish flower-head, that often measures three inches across, with a formidable array of prickly small leaves just below it. In case a would-be pilferer breaks through these lines, however, there is a slight glutinous strip on the outside of the bracts that compose the cup wherein the nectar-filled florets are packed; and here, in sight of Mecca, he meets his death, just as a bird is caught on limed twigs. The pasture thistle, whose range is only from Maine to Delaware, blooms from July to September.
Even gentle Professor Gray hurls anathema at the CANADA THISTLE; "a vile pest" he calls it. As CURSED, CORN, HARD, and CREEPING THISTLE it is variously known here and in Europe, whence it came to overrun our land from Newfoundland to Virginia, westward to Nebraska. By horizontal rootstocks it creeps and forms patches almost impossible to eradicate. The small reddish-purple flower-heads, barely an inch across, usually contain about a hundred florets each. In their tubes the abundant nectar rises high, so that numerous insects, even with the shortest tongues, are able to enjoy it. Not only bees and butterflies, but wasps, flies, and beetles feast diligently. When a floret opens, a quantity of pollen emerges at the upper end of the anther cylinder, pressed up by the growing style. Owing to their slight stickiness and the sharp processes over their entire surface, the pollen grains, which readily cling to the hairs of insects, are transported to the two-branched, hairy stigma of an older floret. But even should insects not visit the flower (and in fine weather they swarm about it), it is marvelously adapted to fertilize itself. Farmers may well despair of exterminating a plant so perfectly equipped in every part; to win life's battles.
"The colour of purple…was, amongst the ancients, typical of royalty. It was a kind of red richly shot with blue, and the dye producing it was attained from a shell found in considerable numbers off the coast of Tyre, and on the shore near the site of that ancient city, great heaps of such shells are still to be found. The production of the true royal purple dye was a very costly affair, and therefore it was often imitated with a mixture of cochineal and indigo…" - J. JAMES TISSOT.
As many so-called purple flowers are more strictly magenta, the reader is referred to the next group if he has not found the flower for which he is in search here. Also to the "White and Greenish" section since many colored flowers show a tendency to revert to the white type from which, doubtless, all were evolved. He should remember that all flowers are more or less variable in shade, according to varying conditions.
"Botany is a sequel of murder and a chronicle of the dead." -JULIAN HAWTHORNE.
"A plant is not to be studied as an absolutely dead thing, but rather as a sentient being…. To measure petals, to count stamens, to describe pistils without reference to their functions, or the why and wherefore of their existence, is to content one's self with husks in the presence of a feast of fatness - to listen to the rattle of dry bones rather than the heavenly harmonies of life. We have reason to be profoundly thankful for the signs to be seen on every side, that the dreary stuff which was called botany in the teaching of the past will soon cease to masquerade in its stolen costume, and that our children and our children's children will study not dried specimens or drier books, but the living things which Nature furnishes in such profusion….
"The reason of this radical change is not far to seek. Since man has learned that the universal brotherhood of life includes himself as the highest link in the chain of organic creation, his interest in all things that live and move and have a being has greatly increased. The movements of the monad now appeal to him in a way that was impossible under the old conceptions. He sees in each of the millions of living forms with which the earth is teeming, the action of many of the laws which are operating in himself; and has learned that to a great extent his welfare is dependent on these seemingly insignificant relations; that in ways undreamed of a century ago they affect human progress." - CLARENCE MOORES WEED.
SESSILE-LEAVED TWISTED-STALK(Streptopus roseus) Lily-of-the-Valley family
Flowers - Dull, purplish pink, 1/2 in. long or less, solitary, on threadlike, curved footstalks longer than the small flower itself, nodding from leaf-axils. Perianth bill-shaped, of 6 spreading segments; stamens 6, 2-horned; style spreading into 3 branches, stigmatic on inner side. Stem: 1 to 2 1/2 ft. high, simple or forked. Leaves: Thin, alternate, green on both sides, many nerved, tapering at end, rounded at base, where they are seated on stem. Fruit: A round, red, many-seeded berry. Preferred Habitat - Moist woods. Flowering Season - May-July. Distribution - North America east and west, southward to Georgia and Oregon.
As we look down on this graceful plant, no blossoms are visible; but if we bend the zig-zagged stem backward, we shall discover the little rosy bells swaying from the base of the leaves on curved footstalks (streptos = twisted, pous = a foot or stalk) very much as the plant's relatives the Solomon's seals grow. In the confident expectation of having its seeds dropped far and wide, it bears showy red berries in August for the birds now wandering through the woods with increased, hungry families.
The CLASPING-LEAVED TWISTED-STALK (S. amplexifolius), which has one or two greenish-white bells nodding from its axils, may be distinguished when not in flower by its leaves, which are hoary - not green - on the under side, or by its oval berry. Indeed most plants living in wet soil have a coating of down on the under sides of their leaves to prevent the pores from clogging with rising vapors.
MOCCASIN FLOWER; PINK, VENUS', or STEMLESS LADY'S SLIPPER(Cypripedium acaule) Orchid family
Flowers - Fragrant, solitary, large, showy, drooping from end of scape, 6 to 12 in. high. Sepals lance-shaped, spreading, greenish purple, 2 in. long or less; petals narrower and longer than sepals. Lip an inflated sac, often over 2 in. long, slit down the middle, and folded inwardly above, pale magenta, veined with darker pink upper part of interior crested with long white hairs. Stamens united with style into unsymmetrical declined column, bearing an anther on either side, and a dilated triangular petal-like sterile stamen above, arching over the broad concave stigma. Leaves: 2, from the base; elliptic, thick, 6 to 8 in. long. Preferred Habitat - Deep, rocky, or sandy woods. Flowering Season - May-June. Distribution - Canada southward to North Carolina, westward to Minnesota and Kentucky.
Because most people cannot forbear picking this exquisite flower that seems too beautiful to be found outside a millionaire's hothouse, it is becoming rarer every year, until the finding of one in the deep forest, where it must now hide, has become the event of a day's walk. Once it was the commonest of the orchids.
"Cross-fertilization," says Darwin, "results in offspring which vanquish the offspring of self-fertilization in the struggle for existence." This has been the motto of the orchid family for ages. No group of plants has taken more elaborate precautions against self-pollination or developed more elaborate and ingenious mechanism to compel insects to transfer their pollen than this.
The fissure down the front of the pink lady's slipper is not so wide but that a bee must use some force to push against its elastic sloping sides and enter the large banquet chamber where he finds generous entertainment secreted among the fine white hairs in the upper part. Presently he has feasted enough. Now one can hear him buzzing about inside, trying to find a way out of the trap. Toward the two little gleams of light through apertures at the end of a passage beyond the nectary hairs, he at length finds his way. Narrower and narrower grows the passage until it would seem as if he could never struggle through; nor can he until his back has rubbed along the sticky, overhanging stigma, which is furnished with minute, rigid, sharply pointed papillae, all directed forward, and placed there for the express purpose of combing out the pollen he has brought from another flower on his back or head. The imported pollen having been safely removed, he still has to struggle on toward freedom through one of the narrow openings, where an anther almost blocks his way.
As he works outward, this anther, drawn downward on its hinge, plasters his back with yellow granular pollen as a parting gift, and away he flies to another lady's slipper to have it combed out by the sticky stigma as described above. The smallest bees can squeeze through the passage without paying toll. To those of the Andrena and Halictus tribe the flower is evidently best adapted. Sometimes the largest bumblebees, either unable or unwilling to get out by the legitimate route, bite their way to liberty. Mutilated sacs are not uncommon. But when unable to get out by fair means, and too bewildered to escape by foul, the large bee must sometimes perish miserably in his gorgeous prison.
SHOWY, GAY, or SPRING ORCHIS(Orchis spectabilis) Orchid family
Flowers - Purplish pink, of deeper and lighter shade, the lower lip white, and thick of texture; from 3 to 6 on a spike; fragrant. Sepals pointed, united, arching above the converging petals, and resembling a hood; lip large, spreading, prolonged into a spur, which is largest at the tip and as long as the twisted footstem. Sterm: 4 to 12 in. high, thick, fleshy, 5-sided. Leaves: 2 large, broadly ovate, glossy green, silvery on under side, rising from a few scales from root. Fruit: A sharply angled capsule, 1 in. long. Preferred Habitat - Rich, moist woods, especially under hemlocks. Flowering Season - April-June. Distribution - From New Brunswick and Ontario southward to our Southern States, westward to Nebraska.
Of the six floral leaves which every orchid, terrestrial or aerial, possesses, one is always peculiar in form, pouch-shaped, or a cornucopia filled with nectar, or a flaunted, fringed banner, or a broad platform for the insect visitors to alight on. Some orchids look to imaginative eyes as if they were masquerading in the disguise of bees, moths, frogs, birds, butterflies. A number of these queer freaks are to be found in Europe. Spring traps, adhesive plasters, and hair-triggers attached to explosive shells of pollen are among the many devices by which orchids compel insects to cross-fertilize them, these flowers as a family showing the most marvelous mechanism adapted to their requirements from insects in the whole floral kingdom. No other blossoms can so well afford to wear magenta, the ugliest shade nature produces, the "lovely rosy purple" of Dutch bulb growers, a color that has an unpleasant effect on not a few American stomachs outside of Hoboken.
But an orchid, from the amazing cleverness of its operations, is attractive under any circumstances to whomever understands it. This earliest member of the family to appear charms the female bumblebee, to whose anatomy it is especially adapted. The males, whose faces are hairy where the females' are bare, and therefore not calculated to retain the sticky pollen masses, are not yet flying when the showy orchis blooms. Bombus Americanorum, which can drain the longest spurs, B. separatus, B. terricola, and, rarely, butterflies as well, have been caught with its pollen masses attached. The bee alights on the projecting lip, pushes her head into the mouth of the corolla, and, as she sips the nectar from the horn of plenty, ruptures by the slight pressure a membrane of the pouch where two sticky buttons, to which two pollen masses are attached, lie imbedded. Instantly after contact these adhere to the round bare spots on her face, the viscid cement hardening before her head is fairly withdrawn. Now the diverging pollen masses, that look like antennae, fall from the perpendicular, by remarkable power of contraction, to a horizontal attitude, that they may be in the precise position to fertilize the stigma of the next flower visited - just as if they possessed a reasoning intelligence! Even after all the pollen has been deposited on the sticky stigmas of various blossoms, stump-like caudicles to which the two little sacs were attached have been found still plastered on a long-suffering bee. But so rich in nectar are the moisture-loving orchids that, to obtain a draught, the sticky plasters which she must carry do not seem too dear a price to pay. In this showy orchis the nectar often rises an eighth of an inch in the tube, and sufficient pressure to cause a rupture will eject it a foot.
ROSE or SWEET POGONIA; SNAKE-MOUTH(Pogonia ophioglossoides) Orchid family
Flowers - Pale rose pink, fragrant, about 1 in. long, usually solitary at end of stem 8 to 15 in. high, and subtended by a leaf-like bract. Sepals and petals equal, oval, about 1/2 in. long, the lip spoon-shaped, crested, and fringed. Column shorter than petals, thick, club-shaped. Anther terminal, attached to back of column, pollen mass in each of its 2 sacs. Stigma a flattened disk below anther. Leaves: 1 to 3, erect, lance-oblong, sometimes one with long footstem from fibrous root. Preferred Habitat - Swamps and low meadows. Flowering Season - June-July. Distribution - Canada to Florida, westward to Kansas.
Rearing its head above the low sedges, often brightened with colonies of the grass pink at the same time, this shy recluse of the swamps woos the passing bee with lovely color, a fragrance like fresh red raspberries, an alluring alighting place all fringed and crested, and with the prospect of hospitable entertainment in the nectary beyond. So in she goes, between the platform and the column overhead, pushing first her head, then brushing her back against the stigma just below the end of the thick column that almost closes the passage. Any powdery pollen she brought on her back from another pogonia must now be brushed off against the sticky stigma. Her feast ended, out she backs. And now a wonderful thing happens. The lid of the anther which is at the end of the column, catching in her shoulders, swings outward on its elastic hinge, releasing a little shower of golden dust, which she must carry on the hairs of her head or back until the sticky stigma of the next pogonia entered kindly wipes it off! This is one of the few orchids whose pollen, usually found in masses, is not united by threads. Without the bee's aid in releasing it from its little box, the lovely species would quickly perish from the face of the earth.
ARETHUSA; INDIAN PINK(Arethusa bulbosa) Orchid family
Flowers - 1 to 2 in. long, bright purple pink, solitary, violet scented, rising from between a pair of small scales at end of smooth scape from 5 to 10 in. high. Lip dropping beneath sepals and petals, broad, rounded, toothed, or fringed, blotched with purple, and with three hairy ridges down its surface. Leaf: Solitary, hidden at first, coming after the flower, but attaining length of 6 in. Root: Bulbous. Fruit: A 6-ribbed capsule, 1 in. long, rarely maturing. Preferred Habitat - Northern bogs and swamps. Flowering Season - May-June. Distribution - From North Carolina and Indiana northward to the Fur Countries.
One flower to a plant, and that one rarely maturing seed; a temptingly beautiful prize which few refrain from carrying home, to have it wither on the way pursued by that more persistent lover than Alpheus, the orchid-hunter who exports the bulbs to European collectors - little wonder this exquisite orchid is rare, and that from certain of those cranberry bogs of Eastern New England, which it formerly brightened with its vivid pink, it has now gone forever. Like Arethusa, the nymph whom Diana changed into a fountain that she might escape from the infatuated river god, Linnaeus fancied this flower a maiden in the midst of a spring bubbling from wet places where presumably none may follow her.
But the bee, our Arethusa's devoted lover, although no villain, still pursues her. He knows that moisture-loving plants secrete the most nectar. When the head of the bee enters the flower to sip, nothing happens; but as he raises his head to depart, it cannot help lifting the lid of the helmet-shaped anther and so letting fall a few soft pellets of pollen on it. Now, after he has drained the next arethusa, his pollen-laden head must rub against the long sticky stigma before it touches the helmet-like anther lid and precipitates another volley of pollen. In some such manner most of our orchids compel insects to work for them in preventing self-fertilization.
Another charming, but much smaller, orchid, that we must don our rubber boots to find where it hides in cool, peaty bogs from Canada and the Northern United States to California, and southward in the Rockies to Arizona, is the CALYPSO (Calypso bulbosa). It is a solitary little flower, standing out from the top of a jointed scape that never rises more than six inches from the solid bulb, hidden in the moss, nor boasts more than one nearly round leaf near its base. The blossom itself suggests one of the lady's slipper orchids, with its rosy purple, narrow, pointed sepals and petals clustered at the top above a large, sac-shaped, whitish lip. The latter is divided into two parts, heavily blotched with cinnamon brown, and woolly with a patch of yellow hairs near the point of the division. May - June.
CALOPOGON; GRASS PINK (Limodorum tuberosum; Calopogon pulchellus of Gray) Orchid family
Flowers - Purplish pink, 1 in. long, 3 to 15 around a long, loose spike. Sepals and petals similar, oval, acute; the lip on upper side of flower is broad at the summit, tapering into a claw, flexible as if hinged, densely bearded on its face with white, yellow, and magenta hairs (Calopogon = beautiful beard). Column below lip (ovary not twisted in this exceptional case); sticky stigma at summit of column, and just below it a 2-celled anther, each cell containing 2 pollen masses, the grain lightly connected by threads. Scape: 1 to 1 1/2 ft. high, slender, naked. Leaf: Solitary, long, grass-like, from a round bulb arising from bulb of previous year. Preferred Habitat - Swamps, cranberry bogs, and low meadows. Flowering Season - June-July. Distribution - Newfoundland to Florida, and westward to the Mississippi.
Fortunately this lovely orchid, one of the most interesting of its highly organized family, is far from rare, and where we find the rose pogonia and other bog-loving relatives growing, the calopogon usually outnumbers them all. Limodorum translated reads meadow-gift; but we find the flower less frequently in grassy places than those who have waded into its favorite haunts could wish.
Owing to the crested lip being oddly situated on the upper part of the flower, which appears to be growing upside down in consequence, one might suppose a visiting insect would not choose to alight on it. The pretty club-shaped, vari-colored hairs, which he may mistake for stamens, and which keep his feet from slipping, irresistibly invite him there, however, when, presto! down drops the fringed lip with startling suddenness. Of course, the bee strikes his back against the column when he falls. Now, there are two slightly upturned little wings on either side of the column, which keep his body from slipping off at either side and necessitate its exit from the end where the stigma smears it with viscid matter. The pressure of the insect on this part starts the pollen masses from their pocket just below; and as the bee slides off the end of the column, the exposed, cobwebby threads to which the pollen grains are attached cling to his sticky body. The sticky substance instantly hardening, the pollen masses, which are drawn out from their pocket as he escapes, are cemented to his abdomen in the precise spot where they must strike against the stigma of the next calopogon he tumbles in; hence cross-fertilization results. What recompense does the bee get for such rough handling? None at all, so far as is known. The flower, which secretes no nectar, is doubtless one of those gay deceivers that Sprengel named "Scheinsaftblumen," only it leads its visitors to look for pollen instead of nectar, on the supposition that the club-shaped hairs on the crests are stamens. The wonder is that the intelligent little bees (a species of Andrenidae), which chiefly are its Victims, have not yet learned to boycott it.